High level of endemism in Haiti’s last remaining forests: a revision of Modisimus (Araneae: Pholcidae) on Hispaniola, using morphology and molecules Author Huber, Bernhard A. Author Fischer, Nadine Author Astrin, Jonas J. text Zoological Journal of the Linnean Society 2010 2010-01-25 158 2 244 299 http://dx.doi.org/10.1111/j.1096-3642.2009.00559.x journal article 10.1111/j.1096-3642.2009.00559.x 0024-4082 5438272 MODISIMUS BACHATA HUBER & FISCHER SP. NOV. ( Figs 55 , 77 , 169–172 , 200 ) Type: Male holotype from near La Ciénaga , (~ 19°03 ′ N , 70°53 ′ W ), La Vega Prov. , Dominican Republic ; ~ 1100 m a.s.l. , path along river, under dry rolled-up leaves on ground, 9 November 2005 ( B.A. Huber ), in ZFMK ( DR 19 a) . Etymology: The species name refers to bachata, a form of music and dance that originated in the Dominican Republic and in which tales of heartbreak and sadness are prevalent. The name is used as a noun in apposition. Diagnosis: Medium-sized species, easily distinguished from congeners by procursus shape ( Fig. 169 ; narrow with dorsodistal projection), and club-shaped hairs on male chelicerae ( Figs 170, 171 ; some hairs on distinct lateral apophyses). Male ( holotype ): Total length, 1.9; carapace width, 0.8. Leg 1: 13.9 (3.3 + 0.3 + 3.5 + 5.3 + 1.5); tibia 2, 2.0; tibia 3, 1.6; tibia 4, 2.1. Tibia 1 L/d: 47. Habitus similar to M. jima sp. nov. (cf. Fig. 19 ), carapace pale ochre-yellow, with dark median line and wider dark median band visible through cuticle; ocular area also darker, clypeus with pair of brown stripes; sternum mostly light brown, lighter medially; legs ochre to light brown, tips of femora and tibiae lighter, very indistinct darker rings subdistally on femora and tibiae; abdomen bluish grey, black spots dorsally and laterally; light brown genital area and plate in front of spinnerets. Ocular area elevated; thoracic furrow distinct. PME–PME, 70 Mm; PME diameter, 105 Mm; PME–ALE, 115 Mm; AME tiny, but with lenses. Sternum wider than long (0.6/0.5), unmodified. Chelicerae with short modified hairs in distinctive pattern, each proximal group on prominent apophysis, distal group barely elevated ( Figs 170, 171 ). Palps as in Figure 169 , coxa with indistinct retrolateral apophysis, femur with proximal flap retrolaterally, and distal apophysis ventrally; procursus very slender, with distinctive distal structures (mostly membranous). Legs with many short vertical hairs on all femora; few spines distally on femur 1, about ten spines in two rows distally on each femur 2; no curved hairs; retrolateral trichobothrium on tibia 1 at 13%; prolateral trichobothrium missing on tibia 1, but present on other tibiae; tarsus 1 with ~25 pseudosegments. Variation: Some males with longer rows of about ten spines each on femur 2, other males without any spines. Tibia 1 in seven other males: 3.2–3.5 (mean 3.4). All males in AMNH appear artificially darkened. Female: In general, similar to male. Tibia 1 in three females : 2.2, 2.2, and 2.3. Epigynum, a very simple plate ( Fig. 55 ); dorsal view as in Figures 77 and 172 . Distribution: Known only from the Ciénaga area, in the central Dominican Republic ( Fig. 200 ). Material examined: Dominican Republic : La Vega Prov. , near La Ciénaga , 1♂ , holotype above; same data, 1♂ , 3♀ and one juvenile ( ZFMK , DR 19 ) ; Parque Nacional A. Bermudez , Ciénaga , 1010 m a.s.l. , tropical evergreen forest, 19 July–2 August 1995 ( S. & J. Peck ), 4♂ and two juveniles #95-33 ( AMNH ) ; same data but 1100 m a.s.l. , 1♂ #95-36 ( AMNH ) ; same data but 1020 m a.s.l. , 1♂ #95-34 ( AMNH ) .