Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa Author GUINOT, DANIÈLE Author TAVARES, MARCOS Author CASTRO, PETER text Zootaxa 2013 2013-05-29 3665 1 1 414 http://dx.doi.org/10.11646/zootaxa.3665.1.1 journal article 10.11646/zootaxa.3665.1.1 1175-5326 5276390 8358B363-BEE3-416D-96CA-8614E38B61D5 Family Pilumnoididae The male gonopore is coxal. The penis emerges from the large gonopore situated on the P5 coxo-sternal condyle in close proximity to the distal extremity of the condyle, more precisely on its anterior margin; the thick and recurved remaining condylar portion articulates on sternite 8 ( Guinot 1979a : fig. 21A; Guinot & Macpherson 1987 : fig. 1) (see Modalities of penis protection : Condylar protection ). The precise relationships of the family remain unknown ( Guinot & Macpherson 1987 ), with, however, larval results suggesting affinities with Carpiliidae ( Luppi et al . 2010 ) .