A late Paleocene fauna from shallow-water chemosynthesis-based ecosystems, Spitsbergen, Svalbard
Author
Hryniewicz, Krzysztof
Author
Amano, Kazutaka
Author
Bitner, Maria Aleksandra
Author
Hagström, Jonas
Author
Kiel, Steffen
Author
Klompmaker, Adiël A.
Author
Mörs, Thomas
Author
Robins, Cristina M.
Author
Kaim, Andrzej
text
Acta Palaeontologica Polonica
2019
2019-02-13
64
1
101
141
http://dx.doi.org/10.4202/app.00554.2018
journal article
10.4202/app.00554.2018
1732-2421
10980900
Genus
Valamunida
Klompmaker and Robins
nov.
ZooBank LSID
:
urn:lsid:zoobank.org:act:
2BA78BB3-34F8-4686-8C5CF70F0416A041
Type
species
:
Valamunida haeggi
Klompmaker and Robins
sp. nov.
; by monotypy, see below.
Etymology
: From the Norse
völva
, an ancient seeress who carried a wand, similar to the rostrum on this galatheoid and -
munida
referring to the family name.
Diagnosis.
—As for the
type
species by monotypy.
Remarks
.—The new genus is placed within the
Munididae
based on a combination of tubercles and spines on the anterior carapace and its lateral margins, the trifid rostrum, and the (bases of) spines at the outer orbital angle. These characters are typical for most
Munididae
, but this combination of characters is not typical for other galatheoid families (
Ahyong et al. 2010
).
Valamunida
Klompmaker and Robins
gen. nov.
is established on the basis of a combination of the tubercular ornamentation across the epibranchial and hepatic regions, more transversely elongate transverse ridges on the posterior carapace, an epigastric region with varied ornamentation, lack of a strong, forwardly-oriented spine on the mesogastric region, and a strongly depressed intestinal region posteriorly.
Valamunida
Klompmaker and Robins
gen. nov.
differs from all 25 known munidid genera. Tubercular ornamentation on the epibranchial and hepatic regions is uncommon in munidids and can only be found in the fossil genera
Juracrista
Robins, Feldmann, and Schweitzer, 2012
and
Protomunida
Beurlen, 1930
, whereas it is also present in modern
Babamunida
Cabezas, Macpherson, and Machordom, 2008
,
Bathymunida
Balss, 1914
,
Hendersonida
Cabezas and Macpherson, 2014
,
Heteronida
Baba and de Saint Laurent, 1996
,
Paramunida
Baba, 1988
,
Plesionida
Baba and de Saint Laurent, 1996
, and
Tasmanida
Ahyong, 2007
. All other genera do not exhibit these previously elaborated features and most of them also lack a strongly depressed posterior part of the intestinal region. From the leftover genera,
Babamunida
,
Juracrista
, and
Protomunida
exhibit only rows of tubercles/spines on the epigastric regions, lacking a much stronger spine. They also lack a strongly depressed posterior part of the intestinal region. Unlike the new genus,
Bathymunida
,
Heteronida
, and
Tasmanida
bear a strong mesogastric spine, as does the
type
species of
Plesionida
.
Plesionida
,
Paramunida
, and
Hendersonida
have a much more granular, spinose to tubercular posterior carapace than
Valamunida
Klompmaker and Robins
gen. nov.
with its transverse ridges.
Paramunida
and
Hendersonida
also lack the depression in the intestinal region.