Order Chiroptera - Family Phyllostomidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 395 426 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Anoura cultrata Handley 1960 Anoura cultrata Handley 1960 , Proc. U. S. Natl. Mus., 112: 463 . Type Locality: Panama , Darién , Río Pucro, Tacarcuna Village, 3,200 ft. ( 1,033 m ); 08°10'N , 77°18'W . Vernacular Names: Handley's Tailless Bat . Synonyms: Anoura brevirostrum Carter 1968 ; Anoura werckleae Starrett 1969 . Distribution: Costa Rica , Panama , Venezuela , Colombia , Ecuador , Peru , Bolivia . Conservation: IUCN 2003 and IUCN / SSC Action Plan (2001) – Lower Risk (lc). Discussion: Includes brevirostrum and werckleae ; see Nagorsen and Tamsitt (1981) and Molinari (1994) . See Tamsitt and Nagorsen (1982) .