Paraneseuthia Franz in the Indian Himalayan Region and Western Palaearctic (Coleoptera, Staphylinidae, Scydmaeninae) Author Jałoszyński, Paweł text Zootaxa 2015 3948 2 287 295 journal article 10.11646/zootaxa.3948.2.9 42729a3f-7efb-44eb-84ca-2bacd77c3f25 1175-5326 243447 ED0782D7-9FFC-4A3B-B521-C418D6C5F459 Paraneseuthia loebli sp. n. ( Figs. 1 , 3–8 ) Type material. Holotype : INDIA (Uttarakhand State): ♂, two labels: " INDE UTTAR PR. / Kumaon, Ranghar / 2250m , 9.X.79 / I. Löbl (7)" [white, printed]; " PARANESEUTHIA / loebli m. / det. P. Jałoszyński, '13 / HOLOTYPUS " [red, printed] ( MHNG ). Paratypes (16 exs): 2 ♀♀, same data as holotype ; 4 ♂♂, 10 ♀♀, same data except for 2400 m (8b), 10.X. [19]79 ( MHNG , cPJ). Each paratype with an additional yellow printed label " PARANESEUTHIA / loebli m. / det. P. Jałoszyński, '15 / PARATYPUS ". FIGURES 1–2. Dorsal habitus of holotype males. Paraneseuthia loebli sp. n. (1); Paraneseuthia meybohmi sp. n. (2). Diagnosis. BL about 1 mm ; body flattened with moderately long and suberect setae; lateral pronotal margins distinctly carinate in posterior half; frons between eyes with shallow transverse impression and pair of shallow lateral pits; protrochanters in males with elongate ventral projection, protibiae in both sexes unmodified; metaventrite in males with longitudinal median carina in posterior half; aedeagus in ventral view with slightly asymmetrical and slightly emarginated apex; parameres strongly recurved in lateral view. Description. BL 1.01–1.05 mm (mean 1.03 mm ). Body of male ( Fig. 1 ) elongate and flattened, light brown, setae yellowish. Head broadest at large, strongly convex and coarsely faceted eyes, HL 0.14–0.15 mm (mean 0.14 mm ), HW 0.20–0.22 mm (mean 0.21 mm ); vertex weakly convex, demarcated from frons by shallow transverse impression extending onto frons between eyes, anterior margin of vertex subtriangular; frons with pair of shallow lateral pits inside transverse impression; supraantennal tubercles small but distinct. Punctures on head dorsum inconspicuous; setae short, sparse and suberect. Antennae short and slender, with indistinctly delimited club composed of antennomeres IX–XI, AnL 0.40–0.42 mm (mean 0.42 mm ), antennomeres I–II elongate, III–VI each about as long as broad, VII–VIII slightly transverse, IX–X distinctly transverse, XI broader than X, about as long as IX–X together, about 1.6× as long as broad. Pronotum subquadrate in dorsal view, with all margins rounded and posterior corners obtuse-angled, broadest near anterior third or near middle (in Fig. 1 pronotum is slightly tilted anteroventrally and its shape differs from that seen in strictly dorsal view); PL 0.27–0.30 mm (mean 0.29 mm ), PW 0.30–0.32 mm (mean 0.31 mm ); pronotal base with shallow transverse impression distinctly deepened at middle and with distinct pair of shallow and small lateral pits. Punctures on pronotal disc fine but distinct, sharply marked and separated by spaces equal to 1.5× diameter of punctures; setae long, sparse and suberect. FIGURES 3–5. Structures of Paraneseuthia loebli sp. n. Pterothorax of female (3) and male (4) in ventral view; mesoventrite of female in ventral view (5). Abbreviations: msvf, mesoventral fovea; mtvc, metaventral carina. FIGURES 6–10. Right fore leg of male in posterior view (6), aedeagus in ventral (7, 9) and lateral (8, 10) views. Paraneseuthia loebli sp. n. (6–8); Paraneseuthia meybohmi sp. n. (9–10). Elytra oval, broadest distinctly anterior to middle and evenly convex; EL 0.57–0.60 mm (mean 0.59 mm ), EW 0.40–0.42 mm (mean 0.42 mm ), EI 1.35–1.50; humeral calli distinct, elytra with short and indistinct basal impressions; apices separately rounded. Punctures on elytra similar to those on pronotum but shallower and with diffused margins; setae long, moderately dense and suberect. Hind wings well developed, about twice as long as elytra. Metaventrite in middle with longitudinal carina in posterior half ( Fig. 4 ; mtvc ). Legs moderately long and slender, protrochanters modified ( Fig. 6 ), with elongate ventral projection, all tibiae nearly straight. Aedeagus ( Figs. 7–8 ) elongate, AeL 0.22 mm ; median lobe in ventral view broadest in basal half and rapidly but only slightly narrowing distally from apical third; apex slightly asymmetrical and slightly emarginate; endophallus with bell-shaped subapical structure; ventral diaphragm circular and located sub-basally; parameres strongly recurved, each with one apical seta. Female. Differs from male in slightly less distinct frontal impression which is less sharply demarcated from vertex, in unmodified protrochanters and metaventrite without carina ( Fig. 3 ); BL 0.97–1.04 mm (mean 1.01 mm ); HL 0.12–0.15 mm (mean 0.14 mm ), HW 0.20–0.22 mm (mean 0.22 mm ), AnL 0.40 mm ; PL 0.27–0.30 mm (mean 0.29 mm ), PW 0.30–0.45 mm (mean 0.33 mm ); EL 0.55–0.60 mm (mean 0.58 mm ), EW 0.40–0.45 mm (mean 0.42 mm ), EI 1.28–1.41. Distribution. India in the Indian Himalayan Region. Etymology. This species is dedicated to Ivan Löbl, the collector of the type series. Remarks. Paraneseuthia loebli differs from all previously known species in a peculiar structure of the mesoventrite ( Figs. 3–5 ), which bears a pair of lateral foveae with setose openings ( Figs. 3–4 ; msvf ). These foveae are directed mesally and their tips are narrowly separated at middle, dorsally to the midline of mesoventral process. In all other characters ( Figs. 3–5 ) this species resembles all congeners.