An illustrated catalogue of the scalpellid barnacles (Crustacea: Cirripedia: Scalpellidae) collected during the HMS “ Challenger ” expedition and deposited in the Natural History Museum, London
Author
Shalaeva, Kate
kshalaeva@googlemail.com
Author
Boxshall, Geoff
kshalaeva@googlemail.com
text
Zootaxa
2014
2014-05-29
3804
1
1
63
journal article
5379
10.11646/zootaxa.3804.1.1
6bf67a5d-ecd7-4f43-b3df-ebcad1333e71
1175-5326
4917932
C1A766F2-D26E-4567-A977-61EF72CBD83A
Trianguloscalpellum regium
(W. Thomson, 1873)
Fig. 33 A, B
Scalpellum regium
Wyville-Thomson, 1873: 347–348
[description only, not figs 1–2 which refer to
Trianguloscalpellum ovale
(
Hoek, 1883
)
].
Scalpellum regium
:
Wyville-Thomson, 1877: 4
, figs 2–3;
Hoek, 1883: 106
, pl. IV, figs 3–5, pl. IX, fig.12, pl. X, figs 1–2; 1884: 10;
Pilsbry, 1907a: 28
, pl. III, fig. 5 [not pl. III, fig. 4 =
T. ovale
(
Hoek, 1883
)
];
Gruvel, 1912: 2
;
1920: 30
, pl. 1, fig. 7;
Zevina, 1970: 263
, figs 10–11; 1973c: 136 [(not
S. regium
, probably
T. gigas
(
Hoek, 1883
)
]; 1976: 1155.
Scalpellum molle
Aurivillius, 1898: 191
;
Gruvel, 1905: 77
;
1920: 29
, pl. V, figs 10–12;
Nilsson-Cantell, 1955: 218
;
Belloc, 1959: 3
;
Zevina, 1976: 1155
.
FIGURE 31.
Trianguloscalpellum moluccanum
(
Hoek, 1883
)
. “Challenger” Stn 195: holotype, left lateral view. Scale bar 10 mm.
FIGURE 32.
Trianguloscalpellum ovale
(
Hoek, 1883
. “Challenger” Stn 61: holotype, right lateral view. Scale bar 10 mm.
non
T. regium:
Zevina, 1981b: 86
;
Foster & Buckeridge, 1995a: 355
, fig. 6K [(
Trianguloscalpellum moluccanum?
(
Hoek,
1883)].
Scalpellum regium latidorsum
Pilsbry, 1907a: 29
.
Arcoscalpellum regium:
Weisbord, 1977: 271
, pl. 31, fig. 1–5.
Trianguloscalpellum regium:
Zevina, 1978b: 1350
;
Shreider, 1994: 159
;
Foster & Buckeridge, 1995b: 167
, figs 2A–D, 3;
Young, 1998b: 28–30
, figs 20a–h, 22; 1999b: 617, fig. 5B,C; 2001a: 741; 2001b: 464, fig. 7; 2004: 26; 2007: 62–63, fig.
72;
Poltarukha & Zevina, 2006a: 158
;
Poltarukha & Zevina, 2006b: 167
, fig. 5;
Poltarukha, 2007: 20
, fig. 4;
Chan
et al.
,
2009: 125–127, figs 106–108;
Chan
et al.
, 2010: 13–14
, figs 1G, 10;
Poltarukha, 2012: 37–38
, fig.11.
Trianguloscalpellum regium regium
:
Zevina, 1981a: 309–310
, fig. 234; Zevina & Shreider, 1992: 40.
Trianguloscalpellum regium latidorsum:
Rosell, 1991: 22
, fig. 1h–i;
Jones
et al.
, 2000: 253
.
FIGURE 33A.
Trianguloscalpellum regium
(W. Thomson, 1873)
. “Challenger” Stn 61: syntype A (left and right lateral view). Scale bar 10 cm.
Material examined.
Two
syntypes
(both partially disintegrated) NHM
UK
2013.1127–1128, Stn 61: Atlantic, Western Central (North Western Atlantic Basin);
5210 m
.
Supplementary descriptions.
Young (1998b)
showed plates only;
Chan
et al.
(2009)
and
Poltarukha (2012)
described hard and soft parts.
Distribution.
Worldwide excluding polar region (see remarks). Known depth range
1507 to 6135 m
.
Remarks.
This species may be attributed to the
Arcoscalpellum michelottianum
group. Numerous specimens have been described from the Atlantic, Pacific and Indian Oceans and this species may represent a complex of cryptic species. Some authors such as
Zevina (1981a)
and
Poltarukha (2012)
consider that
T. regium
is widely distributed across all regions, with the Pacific
T. moluccanum
treated as a synonym, whereas others retain them as separate species, e.g.
Young (1998b
,
2007
).
According to
Hoek (1883)
, “numerous” specimens of the species were found at two “Challenger” stations: 61 and 63. However, only
two specimens
(both from Stn 61) are now in the NHM collection. The
two specimens
, figured and referred as types by
Young (1998b)
, are treated here as
syntypes
, because it can be demonstrated that these were the specimens observed by Wyville-Thomson in his brief description in 1873.
Both specimens are now in different stages of disintegration. One specimen (
syntype
A) (
Fig. 33A
) has the tergum and the apical part of the carina separated (
Fig. 33B
(b)). The other two fragments (tergum and scutum with the upper latus and upper fragment of the rostral latus joined) appear to belong to the second
syntype
(
syntype
B) (
Fig. 33B
(a, c, d)).