Description of five new coral associated Barnacles of the genus Trevathana (Balanomorpha: Pyrgomatidae) in Pacific Waters Author Chan, Benny Kwok Kan Author Chen, Yi-Yang Author Achituv, Yair Author Lin, Hsiu-Chin text Zootaxa 2017 2017-12-11 4363 2 151 202 journal article 31200 10.11646/zootaxa.4363.2.1 fa170e25-9f64-47da-8b18-59d050bf22d0 1175-5326 1098754 6E8B2053-10EC-49EA-8374-D835EACCE01D Trevathana dongshaensis sp. nov. HTTP://zOOBANK.ORG/URN:LSID:zOOBANK.ORG:AcT:BF 022607 -A624-47ED-B90E-21BD032ED670 FIGUReS 2E, F ;17–22 Material examined. HOLOTyPe : ASIZCR000355, DONGSHA ATOLL ( 20°44.949 N , 116°43.301 E ), TAIWAN , MAy 2014 , cOLL. H.N. CHeN, P.C. TSAI , H.C. LIN, ON cORAL Goniastrea minuta VeRON, 2000 . PARATyPeS : ASIZCR000356, DATA SAMe AS ASIZCR000355. ASIZCR000357, DONGSHA ATOLL ( 20°46.767 N , 116°48.402 E ), TAIWAN , MAy 2014 , cOLL. P.C. TSAI , Y.F. TSAO , Y.W. CHANG , ON cORAL Goniastrea minuta . ASIZCR000358, DATA SAMe AS ASIZCR000357. Diagnosis. ScUTUM WITH VeRy BROAD INTeRLOcKING TOOTH, TOOTH HeIGHT ABOUT 3/4 TIMeS HeIGHT Of ARTIcULAR MARGIN. POSTeRIO-LATeRAL SURfAceS Of POSTeRIOR RAMUS Of cIRRUS III BeARING ARRAy Of cONIcAL SPINeS. TeRGUM WITH INWARD TOOTH, PeRPeNDIcULAR TO BASAL MARGIN. Description. SHeLL cOVeReD By cORAL WHeN ALIVe ( FIG. 2E, F ). SHeLL (3.5 MM BD HOLOTyPe ) fLAT, eLLIPTIcAL, PLATeS fUSeD, WHITe (AfTeR cLeANING AND PReSeRVATION), OUTeR SURfAce WITH 17 RADIATING ROWS, UP TO 5 PROjecTIONS IN ROW (17A). CIRcUMfeReNce Of BASe Of SHeLL WITH>30 SHORT PROjecTIONS, eXTeNDING BeyOND OUTLINe ( FIG. 17B ), ORIfIce eLLIPTIcAL, ROSTRO-cARINAL LeNGTH Of ORIfIce NeARLy 1/3 LeNGTH Of TOTAL ROSTRO-cARINAL DIAMeTeR. ScUTUM AND TeRGUM WHITe, SePARATeD ( FIG. 17C–F ). ScUTUM TRANSVeRSALLy eLONGATeD, TeRGAL MARGIN ABOUT 3 TIMeS LONGeR THAN OccLUDeNT MARGIN. OccLUDeNT MARGIN STRAIGHT. EXTeRNAL SURfAce Of ScUTUM WITH STRAIGHT, MeDIAN, cLOSeD fURROW ReAcHING eNTIRe LeNGTH Of ScUTUM ( FIG. 17C ). ARTIcULAR MARGIN Of ScUTUM WITH WIDe INTeRLOcKING TOOTH, TOOTH HeIGHT ABOUT 3/5 HeIGHT Of ARTIcULAR MARGIN ( FIG. 17C ). ADDUcTOR PLATe ABSeNT ( FIG. 17D ). TeRGUM SUBTRIANGULAR ( FIG. 17E, F ), ScUTAL MARGIN cONcAVe, WITH NOTcH; SPUR TRUNcATeD. EXTeRNAL SURfAce WITH SHALLOW, LONGITUDINAL, MeDIAN fURROW ( FIG. 17E ). INNeR SIDe Of ScUTAL MARGIN WITH SeMIcIRcULAR DePReSSION AccOMMODATING INTeRLOcKING TOOTH ( FIG. 17F ). INNeR SURfAce AN INWARD TOOTH LOcATeD ON SPUR, AXIS Of INTeRNAL TOOTH PARALLeL TO AXIS Of SPUR, TIP Of TOOTH eXTeNDING BeyOND BASAL MARGIN Of TeRGUM ( FIG. 17F ). MAXILLA eLLIPTIcAL ( FIG. 18A ), BI-LOBeD, SeRRULATe SeTAe ON eXTeRIOR AND INTeRIOR MARGINS AND DISTAL eNDS Of LOBeS ( FIG. 18B–D ). MAXILLULe WITH cUTTING eDGe STRAIGHT WITHOUT NOTcH, WITH ROW Of 8 SPINeS ( FIG. 18E ), UPPeR AND LOWeR MARGINS WITH SIMPLe SeTAe ( FIG. 18F–H ). MANDIBLe WITH 5 TeeTH, eXcLUDING INfeRIOR ANGLe ( FIG 19A ). SecOND, THIRD AND fOURTH TeeTH BIDeNTATe ( FIG. 19A–D ). FIRST 3 TeeTH OccUPyING ABOUT 4/5 LeNGTH Of cUTTING eDGe, LOWeR MARGIN SHORT WITH feW SeTAe, INfeRIOR ANGLe WITH SeVeRAL SeTAe DISTALLy ( FIG. 19D ). MANDIBULAR PALP OVAL ( FIG. 19E ), BeARING SeRRULATe SeTAe ON eXTeRIOR MARGIN AND DISTALLy ( FIG. 19F ). LABRUM BILOBeD, LOBeS SePARATeD By V-SHAPeD NOTcH ( FIG. 19G ), 3 TeeTH ON LefT SIDe Of NOTcH (19H), NO TeeTH OBSeRVeD ON RIGHT SIDe ( FIG. 19I ). FIGURE 17. Trevathana dongshaensis sp. nov. shell and opercular valves (A) External vieW (ASIZCR000356) and (B) Internal vieW (ASIZCR000355) of shell. (C) External vieW and (D) Internal vieW of left scutum (ASIZCR000355). (E) External vieW (ASIZCR000356) and (F) Internal vieW (ASIZCR000355) of left tergum. Note that for better comparisons of the internal vieW the scutum and tergum Were inclined. Scale bars in µm. FIGURE 18. Trevathana dongshaensis sp. nov. mouth parts. (A) Maxilla, general vieW (ASIZCR000355). (B) Serrulate setae on distal part of maxilla. (C, D) Serrulate setae on interior margin of maxilla. (E) Maxillule, general vieW (ASIZCR000355). (F) Simple setae on upper margin of maxillule. (G, H.) Large, simple setae on cutting edge and loWer margin of maxillule. Scale bars in µm. FIGURE 19. Trevathana dongshaensis sp. nov. mouth parts. (A) Mandible, general vieW (ASIZCR000355). (B) First and bidentate second teeth of mandible. (C) Bidentate third, fourth and simple fifth teeth of mandible. (D) Inferior angle With simple setae of mandible. (E) Mandibular palp (ASIZCR000355). (F) Serrulate setae on exterior margin of mandibular palp. (G) Labrum (ASIZCR000355). (H, I) Teeth on crest of labrum. Scale bars in µm. FIGURE 20. Trevathana dongshaensis sp. nov. cirri I–II (ASIZCR000355). (A) Cirrus I. (B) Serrulate setae on apex of posterior ramus apex of cirrus I. (C) Serrulate setae on apex of anterior ramus of cirrus I. (D) Serrulate setae on apex of anterior ramus of cirrus I. (E) Cirrus II. (F) Serrulate setae and conical spines on posterior ramus of cirrus II. (G) Fan-shaped denticles on posterior ramus of cirrus II (indicated by White arroWs). (H) Serrulate setae on anterior ramus of cirrus II. Scale bars in µm. FIGURE 21. Trevathana dongshaensis sp. nov. cirrus III–IV. (ASIZCR000355) (A) Cirrus III. (B) Posterior rami of cirrus III. (C) Anterior ramus of cirrus III. (D) Conical spines on apex of posterior ramus of cirrus III (indicated by White arroWs). (E) Serrulate setae on posterior ramus of cirrus III. (F) Cirrus IV. (G) Intermediate segment With 4 pairs of serrulate setae on cirrus IV. (H) Serrulate setae on apex of rami of cirrus IV. Scale bars in µm. FIGURE 22. Trevathana dongshaensis sp. nov. cirri V–VI and penis. (ASIZCR000355) (A) Cirrus V (B) Intermediate segment With 4 pairs of serrulate setae on cirrus V. (C) Distal segment of cirrus VI shoWing serrulate setae. (D) Cirrus VI (E) Intermediate segment With 4 pairs of serrulate setae on cirrus VI. (F) Distal segment of cirrus VI shoWing serrulate setae. (G) Penis (H). Basidorsal piont of penis. (I) Apex of penis. Scale bars in µm. CIRRUS I WITH RAMI SUBeqUAL; ANTeRIOR RAMUS LONG, SLeNDeR, 8 SeGMeNTeD ( FIG. 20A ); POSTeRIOR RAMUS 5 SeGMeNTeD, BOTH RAMI BeARING SeRRULATe SeTAe ( FIG. 20B–D ). CIRRUS II WITH RAMI SLIGHTLy UNeqUAL; ANTeRIOR RAMUS 9- SeGMeNTeD, POSTeRIOR RAMUS 6-SeGMeNTeD ( FIG. 20E ), BeARING SeRRULATe SeTAe ( FIG. 20F, H ); ANTeRIOR Of POSTeRIOR RAMUS BeARING cOMB STRUcTUReS ( FIG. 20G ). CIRRUS III WITH ANTeRIOR AND POSTeRIOR RAMI SIMILAR LeNGTH, 8 SeGMeNTeD ( FIG. 21A ), BeARING SeRRULATe SeTAe ( FIG. 21C–E ); POSTeRIO-LATeRAL SIDe Of POSTeRIOR RAMUS BeARING cONIcAL SPINeS ALONG 3 DISTAL SeGMeNTS ( FIG. 21 , B, D), WITH SeRRULATe SeTAe ( FIG. 21E ). CIRRI IV–VI LONG, SLeNDeR RAMI Of SIMILAR LeNGTHS; cIRRUS IV, ANTeRIOR RAMUS18 SeGMeNTeD, POSTeRIOR RAMUS 19 SeGMeNTeD; cIRRUS V (20, 20); cIRRUS VI (21, 21) ( FIGS. 21F , 22A, D ); ALL RAMI BeARING SeRRULATe AND SIMPLe SeTAe ( FIGS. 21G, H , 22B, C, E, F ); INTeRMeDIATe SeGMeNTS Of RAMI Of CIRRI IV AND V BeARING 3 PAIRS Of LONG, SeRRULATe SeTAe AND 1 PAIR Of SIMPLe SeTAe ( FIGS. 21G, H , 22B, E, F ), DISTAL PAIR LONGeST, PROXIMAL PAIR SHORTeST. PeNIS ANNULATeD, WITH ScATTeReD SHORT SIMPLe-TyPe SeTAe ( FIG. 22G ). PeDIceL WITH BLUNT BASIDORSAL POINT ( FIG. 22H ), APeX BeARING SHORT, SIMPLe SeTAe ( FIG. 22I ). HABITAT. ON cORAL Goniastrea minuta Distribution. AT PReSeNT, ONLy RecORDeD IN THe DONGSHA ATOLL, TAIWAN . Etymology. Trevathana dongshaensis IS NAMeD fROM ITS TyPe LOcALITy, DONGSHA ATOLL, TAIWAN . Remarks. Trevathana dongshaensis HAS AVeRy BROAD, INTeRLOcKING TOOTH ON ScUTUM, WHIcH IS SIMILAR TO THAT fOUND IN T. paulayi . Trevathana dongshaensis DIffeRS fROM T. paulayi By HAVING THe INTeRNAL TOOTH Of THe TeRGUM eXTeNDING BeyOND THe BASAL MARGIN.