A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico Author Luna-Cozar, Jesús jlcozar@yahoo.com.mx Author Anderson, Robert S. jlcozar@yahoo.com.mx Author Jones, Robert W. Author León-Cortés, Jorge L. text Zootaxa 2014 2014-04-15 3788 1 1 63 journal article 5629 10.11646/zootaxa.3788.1 80d33caa-2379-4f5e-ba98-6d1cc29a7906 1175-5326 4913761 7484832A-025B-41FA-9696-DF7531C01AA4 Tylodinus mutabilis species group (2) Tylodinus mutabilis new species ; México ( Chiapas : La Concordia, Villacorzo) Tylodinus parvus new species ; México ( Chiapas : Trinitaria) Tylodinus nodulosus species group (3) Tylodinus andersoni new species ; México ( Chiapas : Ángel Albino Corzo) Tylodinus nodulosus (Boheman) 1837 ; México Tylodinus zilchi Kuschel 1956 ; El Salvador , Honduras Tylodinus pusillus species group (2) Tylodinus porvenirensis new species ; México ( Chiapas : El Porvenir) Tylodinus pusillus new species ; México ( Chiapas : Ángel Albino Corzo, La Concordia, Unión Juárez) Tylodinus spiniventris species group (2) Tylodinus lum new species ; México ( Chiapas : Coapilla, San Cristóbal de las Casas) Tylodinus spiniventris new species ; México ( Chiapas : Huixtán, San Cristóbal de las Casas, Pueblo Nuevo Solistahuacán). Incertae sedis (13) Tylodinus complicatus new species ; México ( Chiapas : Bochil, Coapilla, El Bosque, Rayón, Tapalapa) Tylodinus dominicus new species ; México ( Chiapas : Villacorzo) Tylodinus noctis new species ; México ( Chiapas : Coapilla, Rayón) Tylodinus rufus new species ; México ( Chiapas : Chalchihuitán, Coapilla, Huixtán, San Cristóbal de las Casa, San Juan Chamula, Tapalapa) Tylodinus branstetteri new species ; México ( Chiapas : La Concordia) Tylodinus kuscheli new species ; México ( Chiapas : Villacorzo) Tylodinus pappi new species ; México ( Chiapas : Unión Juárez) Tylodinus gibbosus new species ; México ( Chiapas : Pueblo Nuevo Solistahuacán) Tylodinus immundus new species ; México ( Chiapas : San Cristóbal de las Casas) Tylodinus intzin new species ; México ( Chiapas : Bochil, Chalchihuitán, El Bosque, Pueblo Nuevo Solistahuacán, Rayón, Tenejapa, Trinitaria) Tylodinus elongatus new species ; México ( Chiapas : Angel Albino Corzo) Tylodinus kissingeri new species ; México ( Chiapas : Chalchihuitán, Tapalapa) Tylodinus pinguis new species ; México ( Chiapas : Ángel Albino Corzo, La Concordia) Key to species and species groups of Tylodinus Champion from Chiapas 1 Elytral declivity with laminate or elongate tubercles ( Figures 29, 31, 33 , 96–97 ).................................... 2 - Elytral declivity with rounded tubercles ( Figures 34–95 ).......................................................3 2(1) Pronotal disc at most slightly longitudinally impressed; elytra disc with rounded tubercles ( Figures. 96–97 ).................................................................................................. T. pinguis Luna-Cozar - Pronotal disc deeply excavate medially; elytra disc with laminate tubercles (especially that at base of I2, Figures 28–33 ).................................................................................... T. nodulosus species group 3(1) Pronotal disc without impressions; base of elytra declivity with a transverse fringe of white scales ( Figures 40–51 )........................................................................................ T. buchanani species group - Pronotal disc with a longitudinal impression; elytra declivity without white scales ( Figures 34–39 , 52–95 )............... 4 4(3) Elytra black with dark red coloration at base, along interval 2 and on elytral declivity from interval 1 to 5 ( Figures 64–67 , 74–79 ).............................................................................................. 5 - Elytra black ( Figures 52–63 , 68–73 , 80–95 )................................................................. 8 5(4) Hind tibiae with an external tooth ( Figure 26 )............................................................... 6 - Hind tibiae without an external tooth...................................................................... 7 6(5) Metasternal tooth absent; premucro present; with a patch of yellow scales between eyes.......... T. dominicus Luna-Cozar - Metasternal tooth present ( Figure 19 ); premucro absent; scales between eyes similar to scales on head and rostrum......................................................................................... T. mutabilis species group 7(5) Pronotum red ( Figures 76–77 )............................................................ T. rufus Luna-Cozar - Pronotum black ( Figures 74–75 )..................................................... T. complicatus Luna-Cozar 8(4) Front and middle tibiae (less so in female) very deeply preapically excavate ( Fig. 27 ); pronotum and rostrum rugose ( Figures 34–39 )........................................................................... T. cavicrus species group - Front and middle tibiae not deeply excavate, generally straight or slightly curved to sinuate; pronotum and rostrum not rugos. .................................................................................................... 9 9(8) Hind tibiae with an external tooth ( Figure 26 )........................................ T. canaliculatus species group - Hind tibiae without an external tooth..................................................................... 10 10(9) Body very elongate, 2.7 x as long as wide; abdomen longer, as long as the distance from the apex of head to the posterior edge of metasternum ( Figures 72–73 )....................................................... T. elongatus Luna-Cozar - Body not so elongate; abdomen shorter than the length from head to posterior edge of metasternum................... 11 11(10) Body length longer than 3.5 mm .........................................................................12 - Body length 3.5 mm or less............................................................................. 14 12 (11) Median section of ventrite 2 glabrous; tubercles on base of elytral declivity similar in size ( Figures 60–63 ).............................................................................................. T. spiniventris species group - Median section of ventrite 2 with a transverse line of scales along midline, occasionally interrupted medially ( Figures 23–24 ); base of elytral declivity with tubercle on interval 4 longer than tubercle on interval 2............................... 13 13(12) Pronotum in dorsal view with very dense whitish scales on lateral sides; ventrite 1 with scales only on anterior margin, median section glabrous ( Figures 94–95 )...................................................... T. kissingeri Luna-Cozar - Pronotum in dorsal view with very dense yellow to light brown scales on lateral sides; ventrite 1 squamous ( Figures 82–83 ).................................................................................... T. kuscheli Luna-Cozar 14(11) Elytra strongly convex in lateral view; body completely clothed with gray and whitish scales ( Figures 92–93 )............................................................................................... T. gibbosus Luna-Cozar - Elytra not strongly convex ( Figures 87, 89, 91 ); body clothed with dark brown scales, yellow or light brown, if whitish scales are present, then is in patches or scattered and intermixed..................................................... 15 15(14) Body clothed with dark brown or black scales with whitish scales denser along lateral margins of pronotal disc and tubercles in front of elytral declivity ( Figures 52–55 )............................................. T. coapillensis species group - Body without whitish scales............................................................................ 16 16(15) Base of elytral declivity without tubercles on interval 2 ( Figures 90–91 ).......................... T. intzin Luna-Cozar - Base of elytral declivity with a tubercle on interval 2 ( Figures 68–71 , 85, 87 )..................................... 17 17(16) Ventrite 2 squamous................................................................ T. immundus Luna-Cozar - Ventrite 2 glabrous in median section or with a transverse line of scales along midline.............................. 18 18(17) Ventrite 1 squamous................................................................... T. pappi Luna-Cozar - Ventrite 1 with scales only on anterior margin, median section glabrous ( Figures 20, 25 )............................ 19 19(18) Punctures on pronotal disc uniformly dense at the base, progressively more widely spaced, becoming smaller and shallower at apex ( Figures 14 ); base of elytral declivity with tubercle on interval 2 as long as tubercle on interval 4; dermis shiny, metasternum concave..................................................................... .. T. pusillus species group - Punctures on pronotal disc dense to contiguous on sides, progressively more widely spaced and less conspicuous in the median, anterior and posterior sections ( Figure 15 ); base of elytral declivity with tubercle on interval 2 longer or smaller than tubercle on interval 4.................................................................................. 20 20(19) Base of elytral declivity with tubercle on interval 2 longer than tubercle on interval 4; coloration of body scales predominantly light brown ( Figures 86–87 )........................................................ T. branstetteri Luna-Cozar - Base of elytral declivity with tubercle on interval 2 smaller than tubercle on interval 4; coloration of body scales predominantly dark brown ( Figures 80–81 )............................................................. T. noctis Luna-Cozar