A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico
Author
Luna-Cozar, Jesús
jlcozar@yahoo.com.mx
Author
Anderson, Robert S.
jlcozar@yahoo.com.mx
Author
Jones, Robert W.
Author
León-Cortés, Jorge L.
text
Zootaxa
2014
2014-04-15
3788
1
1
63
journal article
5629
10.11646/zootaxa.3788.1
80d33caa-2379-4f5e-ba98-6d1cc29a7906
1175-5326
4913761
7484832A-025B-41FA-9696-DF7531C01AA4
12.
Tylodinus variabilis
Luna-Cozar
,
sp. nov.
(
Figures 2
,
50–51
,
107
,
132
)
Diagnosis.
Length male, 3.4–4.0 mm, female, 3.2–4.0 mm. Width male,
1.6–1.7 mm
, female 1.5–2.0 mm. As for
T. buchanani
with the follow exceptions: body clothed with short scales, light scales on vertex and pronotum frequently whitish, but in some specimens light brown. Head with overlapping scales on vertex; front moderately concave. Elytra with basal margin straight (
Figure 2
), stria 10 complete, surface granulate, tubercles similar to
T. buchanani
. Metasternal tooth absent. Abdomen squamous, scales setigerous, ventrite 1 convex; ventrite 2 longer than 3–4 combined, convex. Hind femora length almost reaching elytral apex, front tibiae sinuate, middle and hind tibiae subparallel, premucro very small, hind tibiae with apical ¼ curved. Male genitalia (
Fig. 107
) in lateral view slender; in ventral view with lateral sides concave, subparallel; apical process in lateral view weakly curved. Female genitalia with tergite eight trapezoidal, apex slightly curved, sternite eight oblong, median membranous area subapical; spermatheca shallowly curved, ramus long and slender, nodulus slender.
Geographic distribution
.
México
(
Chiapas
).
Habitat and elevation.
Specimens were collected from leaf litter in high elevation cloud forest between
2300–2750 m
.
Derivation of specific name.
Name derived from the Latin word
variabilis
= variable.
Material examined
.
Total
95 males
,
90 females
.
Holotype
male (
CMNC
):
México
:
Chiapas
,
San Cristóbal
de las
Casas
,
Cerro Huitepec
(pico)
5 km
W San Cristóbal
, elev.
2750 m
(
16°44'33''N
,
92°41'38.4''W
),
15.ix.1991
, coll.
R
.
S. Anderson.
Paratypes
:
México
:
Chiapas
,
Huixtán
,
15 km
E San Cristóbal
, elev.
2500 m
(
16°44'49.6''N
,
92°29'24.22''W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-24)
—
1 female
(
CMNC
); same locality (
16.74717°N
92.49017°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-48)
—
1 female
(
CMNC
); same locality (
16.74698°N
92.49013°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-16)
—
1 male
(
CMNC
);
Bazom
, elev.
2450 m
(
16°44'18.96''N
,
92°29'18.24''W
),
9.vii.2003
, coll.
R
.
S. Anderson
—
1 male
(
CMNC
);
San Cristobal
de las
Casas Airport
, elev.
2350 m
(
16°41'25.08''N
,
92°31'48''W
),
25.vii.2005
, coll.
R
.
S. Anderson
—
1 female
(
CMNC
);
San Cristóbal
de las
Casas
,
Cerro Huitepec
(pico)
5 km
W San Cristóbal
, elev.
2750 m
(
16°44'33''N
,
92°41'38.4''W
),
15.ix.1991
, coll.
R
.
S.
Anderson
—
6 males
,
6 females
(
CMNC
); same locality,
18.ix.1991
, coll.
R
.
S. Anderson
—
4 males
,
2 females
(
CMNC
);
Cerro Huitepec
5 km
W San Cristóbal
, elev.
2650 m
(
16°44'37''N
,
92°41'23.3''W
),
17.ix.1992
, coll.
R
.
S. Anderson
—
2 males
,
5 females
(
BMNH
,
CMNC
);
Corazón de María
, elev.
2449 m
(
16°41'7.4''N
,
92°32'11.4''W
),
8.vi.2006
, coll.
J. L. Cozar
—
1 male
(
ECOS
);
Huitepec
, elev.
2466 m
(
16°44'54.2''N
,
92°41'6.4''W
),
5.v.2006
, coll.
J. L. Cozar
—
2 males
,
2 females
(
ECOS
); same locality,
15.vi.2006
, coll.
J. L. Cozar
—
1 male
(
ECOS
); same locality,
3.vii.2006
, coll.
J. L. Cozar
—
2 males
(
ECOS
); same locality (
16.75181°N
92.68270°W
), elev.
2480 m
,
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-01)
—
2 males
,
1 female
(
CWOB
); same locality (
16.75104°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-18)
—
1 male
,
1 female
(
CMNC
); same locality (
16.75109°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-17)
—
1 female
(
CMNC
); same locality (
16.75131°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-12)
—
1 female
(
CMNC
); same locality (
16.75176°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-02)
—
3 males
,
1 female
(
CMNC
); same locality (
16.75163°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-05)
—
3 males
,
5 females
(
CMNC
,
ECOS
); same locality (
16.74689°N
92.48990°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-1-01)
—
1 male
(
CMNC
); same locality (
16.75018°N
92.68280°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-39)
—
2 males
(
CMNC
); same locality (
16.75018°N
92.68284°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-40)
—
1 male
(
CMNC
);
Huitepec Ocotal
,
Cerro La Cruz
, elev.
2970 m
(
16°44'31.2''N
,
92°41'37.7''W
),
12.vi.2006
, coll.
J. L. Cozar
—
3 males
,
2 females
(
ECOS
);
Huitepec
,
5 km
W San Cristóbal
, elev.
2750 m
(
16°44'33''N
,
92°41'38.4''W
),
14-25.ix.1992
, coll.
R
.
S.
Anderson
—
2 females
(
CMNC
); same locality,
14.ix.1992
, coll.
R
.
S. Anderson
—
5 males
,
2 females
(
CMNC
);
Huitepec, S
.
Cristóbal
, elev.
2480 m
(
16.75041°N
92.68270°W
),
29.v.2001
, coll.
LLAMA
(Wa-A-05-2-32)
—
1 male
,
1 female
(
CMNC
); same locality (
16.75032°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-34)
—
2 females
(
ECOS
); same locality (
16.75045°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-31)
—
1 male
,
2 females
(
CMNC
); same locality (
16.75163°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-05)
—
1 female
(
CMNC
); same locality (
16.75050°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-30)
—
1 male
,
1 female
(
ECOS
); same locality (
16.75032°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-34)
—
1 female
(
CMNC
); same locality (
16.75077°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-24)
—
1 female
(
CMNC
); same locality (
16.75018°N
92.68298°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-43)
—
1 female
(
CMNC
); same locality (
16.75018°N
92.68303°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-44)
—
1 female
(
CMNC
); same locality (
16.75018°N
92.68308°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-45)
—
1 male
(
CWOB
); same locality (
16.75018°N
92.68317°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-47)
—
1 male
(
CMNC
); same locality (
16.75018°N
92.68327°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-49)
—
1 female
(
CMNC
); same locality (
16.75054°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-29)
—
1 male
(
CMNC
); same locality (
16.75082°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-23)
—
1 male
(
CMNC
);
Los Alcanfores
, elev.
2450 m
(
16°44'12.47''N
,
92°40'46.93''W
),
25.iv.2006
, coll.
M. Girón
—
1 male
(
ECOS
);
Reserva Huitepec
, elev.
2400 m
(
16°44'12.5''N
,
92°40'46.9''W
),
18.xi.1993
, coll.
R
.
Jones
—
4 males
,
1 female
(
ECOS
,
UAQM
); same locality,
18.xi.1993
, coll.
R
.
Jones
—
1 male
(
ECOS
); same locality,
16.ii.1994
, coll.
R
.
Jones
—
1 male
(
ECOS
); same locality,
16.ii.1994
, coll.
R
.
Jones
—
2 males
,
2 females
(
ECOS
); same locality,
7.iv.1994
, coll.
R
.
Jones
—
1 females
(
ECOS
); same locality,
27.v.1994
, coll. B. Gómez-G y
O. L.
Gómez-N—
2 males
(
ECOS
); same locality,
25.viii.1994
, coll. B. Gómez-G—
2 males
(
ECOS
); same locality,
26.x.1994
, coll. B. Gómez-G—
1 male
,
1 female
(
ECOS
); same locality,
24.i.1995
, coll.
R
.
Jones
y B. Gómez-G—
1 female
(
ECOS
); same locality,
24.i.1995
, coll.
R
.
Jones
—
1 male
(
ECOS
); same locality,
24.i.1995
, coll.
R
.
Jones-W y B. Gómez-G—
1 male
(
ECOS
); same locality,
27.iv.1995
, coll. B. Gómez-G y M. Girón-I—
1 female
(
ECOS
); same locality,
14.vi.1995
, coll. B. Gómez- G, M. Girón-I y A. Méndoza-R—
1 male
(
ECOS
); same locality,
27.vi.1995
, coll. B. Gómez-G y M. Girón-I—
1 female
(
ECOS
); same locality,
5.vii.1995
, coll.
M. Girón
—
1 male
(
UAQM
); same locality,
5.vii.1995
, coll.
R
.
Jones
—
1 male
(
UAQM
); same locality,
10.vii.1995
, coll.
R
.
Jones
—
1 female
(
UAQM
); same locality,
25.vii.1995
, coll.
R
.
W.
Jones
—
4 males
,
3 females
(
ECOS
,
UAQM
); same locality,
1.viii.1995
, coll.
A. Mendoza
y
M. Girón
—
1 male
(
UAQM
); same locality,
15.viii.1995
, coll.
M. Girón
—
1 female
(
UAQM
); same locality,
3.xi.1995
, coll.
B. Gómez
y
M. Girón
—
1 male
,
2 females
(
UAQM
); same locality,
14.xi.1995
, coll.
B. Gómez
,
M. Girón
—
1 male
,
1 female
(
UAQM
); same locality,
5.xii.1995
, coll.
B. Gómez
y
M. Girón
—
1 male
,
2 females
(
UAQM
); same locality,
6.ii.1996
, coll. L. Ruiz-M y M. Girón-I—
3 males
,
4 females
(
ECOS
,
UAQM
); same locality,
14.ii.1996
, coll. L. Ruiz-M y M. Girón-I—
1 male
(
ECOS
); same locality,
14.ii.1996
, coll. L. Ruiz-M y M. Girón-I—
2 males
,
2 females
(
UAQM
); same locality,
7.vii.2003
, coll.
R
.
S.
Anderson
—
1 female
(
CMNC
); same locality,
12.ii.2004
, coll.
J. L. Cozar
,
M. Girón
—
1 female
(
ECOS
); same locality,
9.vi.2004
, coll.
J. L. Cozar
,
M. Girón
—
1 male
,
1 female
(
ECOS
); same locality,
11.vi.2004
, coll.
J. L. Cozar
—
1 female
(
ECOS
); same locality,
2.vii.2004
, coll.
J. L. Cozar
—
1 female
(
ECOS
); same locality,
12.vii.2004
, coll.
J. L. Cozar
—
1 female
(
ECOS
); same locality,
13.vii.2004
, coll.
M. Girón
—
1 male
(
ECOS
); same locality,
15.vii.2005
, coll.
R
.
S.
Anderson
—
1 male
(
CMNC
); same locality,
9.vi.2006
, coll.
U. Caballero
—
1 female
(
ECOS
); same locality,
12.vi.2006
, coll.
J. L. Cozar
—
1 female
(
ECOS
); same locality,
17.vi.2006
, coll.
U. Caballero
—
3 females
(
ECOS
); same locality,
11.vii.2007
, coll.
J. Longino
—
1 female
(
CMNC
); same locality,
9.vii.2007
, coll.
R
.
S.
Anderson
—
4 males
,
4 females
(
CMNC
); same locality,
9.vii.2007
, coll.
M. G. Branstetter
—
2 females
,
1 male
(
CMNC
); same locality,
11.vii.2007
, coll.
R
.
S. Anderson
—
3 males
,
2 females
(
CMNC
);
Zoológico
, elev.
2379 m
(
16°43'5.5''N
,
92°42'9.7''W
),
6.vi.2006
, coll.
J. L. Cozar
—
1 male
(
ECOS
);
San
Juan Chamula, Cerro Tzontehuitz, elev.
2800 m
(
16°49'10.2''N
,
92°35'3.8''W
),
15.vii.1994
, coll.
R
.
Jones
—
1 male
(
UAQM
)
.