Eleven new species of Amblypsilopus Bigot (Diptera: Dolichopodidae: Sciapodinae) and a key to the species of Madagascar and adjacent islands Author Grichanov, Igor Ya. text European Journal of Taxonomy 2021 2021-06-18 755 1 47 87 http://dx.doi.org/10.5852/ejt.2021.755.1399 journal article 6004 10.5852/ejt.2021.755.1399 d7d8ccff-7a58-4fc8-9eec-c2588b457d56 2118-9773 4983303 4C7C9B68-4026-45F9-98C4-430FE018119F Amblypsilopus analamazaotra sp. nov. urn:lsid:zoobank.org:act: 5AEAE000-F228-4D8D-B537-887690037C13 Fig. 7 Diagnosis The new species is very close to A. fianarantsoa sp. nov. , differing distinctly in its longer fore tarsus with segment 4 black and simple, segment 2 distinctly shorter than segment 3. Amblypsilopus fianarantsoa sp. nov. differs from the new species in the fore tarsomere 4 yellow, flattened and widened, with posterior comb of 8–9 long flattened setae, segment 2 as long as segment 3. The fore basitarsus is 2–3 times as long as fore tibia, and the last segment of the fore tarsus is white in both species. Etymology The species is named after the Analamazaotra Special Reserve, a wildlife reserve of Madagascar , which is part of Andasibe-Mantadia National Park. The reserve is situated in the region Alaotra-Mangoro, close to Moramanga and Andasibe. Material examined Holotype MADAGASCAR • ♂ ; Andasibe , Analamazaotra Forest ; 18°46ʹ S , 48°24ʹ E ; 950 m a.s.l. ; 31 Oct.–4 Nov. 2007 ; A. Freidberg leg.; SMNHTAU . Paratype MADAGASCAR • 1 ♂ ; same collection data as for holotype; SMNHTAU . Description Male ( Fig. 7A ) MEASUREMENTS. Body length 4.7 mm ; antenna length 1 mm ; wing length 3.6 mm ; wing width 1 mm . HEAD ( Fig. 7B ). Frons greenish black, white pollinose; strong anterior vertical and postvertical bristles; upper postocular setae black, short; lateral postocular setae white, uniserial; ventral postcranium covered with irregular white hairs; face densely silvery-white pollinose, narrow, under antennae 1.5 times as wide as postpedicel, at clypeus almost obliterated; clypeus small, slightly bulging; antenna ( Fig. 7C ) black, 2.2 times as long as height of head; pedicel with short bristles, with one long dorsal bristle; postpedicel rounded, as long as high, with short hairs; arista-like stylus dorsal, microscopically haired; length (mm) of scape, pedicel, postpedicel, stylus (segments 1 and 2), 0.06/0.06/0.09/0.03/0.71; proboscis orange; palpus blackish, with white and brown hairs. THORAX. Mesonotum and scutellum metallic blue-green, weakly pollinose; pleura bronze-green, grey pollinose; metepimeron yellow; 6 dorsocentral bristles gradually decreasing in size anteriorly with strongest posterior pair; acrostichals short, biserial; scutellum with 2 strong bristles and 2 minute hairs laterally. LEGS. Mostly yellow; coxae yellow, but mid coxa orange at base; fore and mid tarsi brownish to brown; segment 5 of fore tarsus silvery white ( Fig. 7D ); hind femur brown on distal half dorsally; hind tibia brownish; hind tarsus brown to black; fore and mid coxae with dark hairs and 2 black subapical setae; hind coxa with 1 black seta at middle; femora without strong or long setae; fore tibia without conspicuous setae; fore basitarsus long, with 3–4 short ventral setae on distal half; mid tibia with 1–2 short anterodorsals and 2 short posterodorsals, without conspicuous ventral setae; mid and hind tarsi simple, cylindrical; hind tibia with several short dorsal and ventral setae; femur, tibia and tarsomere (from first to fifth) length ratio (mm): fore leg: 1.11/1.1/2.92/0.26/0.48/0.32/0.24, mid leg: 1.19/2.14/1.64/0.5/0.31/0.2/0.14, hind leg: 1.57/2.64/1.05/0.57/0.26/0.18/0.14. Fig. 7. Amblypsilopus analamazaotra sp. nov. A–E . Holotype, ♂ (SMNHTAU). F–G . Paratype, ♂ (SMNHTAU). A . Habitus. B . Head. C . Antenna. D . Fore tarsomeres 2–5. E . Wing. F . Hypopygium, left lateral view, reflected light. G . Distoventral appendages of hypopygium, transmitted light. WING ( Fig. 7E ). Widened distally, almost hyaline, veins brown; R 4+5 gently curved to M 1 in apical third; M 1+2 straight in basal half, slightly convex anteriad in middle part; M 1 with strong elbow, forming right angle with M 1+2 ; ratio of parts of costa between R 2+3 and R 4+5 to those between R 4+5 and M 1 , 4/1; crossvein dm-m straight; ratio of crossvein dm-m to apical part of M (fork-handle) to apical part of M 4 , 30/78/21; anal vein and lobe reduced; anal angle absent; lower calypter brownish, with fine yellow cilia; halter yellow with brown knob. 1+2 ABDOMEN. Thin and long, mostly shining dark brown, with short black hairs and marginal setae; segment 1 mostly yellow, with white and dark hairs; pregenital segments combined nearly three times as long as mesonotum; hypopygium ( Fig. 7F ) brown, with brown appendages; cercus brown, bifurcated from base; dorsal arm as long as epandrium, filiform, slightly widened at base, sparsely covered with long light setae along entire length decreasing in length distally, densely covered with hairs at base; ventral arm of cercus S-shaped, narrow, thickened at base and at apex, with two thick apical setae, one very long curved seta at base of small apicodorsal process covered with 5 short and long hairs ( Fig. 7G ); ventral arms of cerci fused almost to apex; surstylus projected, bilobed; dorsal arm of surstylus leaf-like, broad, with deep distal emargination; ventral arm of surstylus trilobed, with apical setae on each lobe, with oval leaf-like seta on outer lobe, with short leaf-like triangular seta and tuft of short hairs at apex of median lobe; epandrial lobe projected, trilobed; each lobe narrow, with pair of long or short flattened setae at apex; outer lobe with 1 simple and 1 hooked seta ( Fig. 7G ). Female Unknown.