Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae) Author Semenyuk, Irina Author Golovatch, Sergei I. Author Wesener, Thomas text Zootaxa 2018 2018-08-16 4459 3 535 550 journal article 29010 10.11646/zootaxa.4459.3.7 81db55ed-b19e-42e9-930e-f9697834cc1b 1175-5326 1458879 8CA301B7-A24A-4CB1-8231-FF0BAFF175DA Key to species of Sphaerobelum (based mainly on male characters): 1. Anterior telopod with 3 telopoditomeres................................................................... 2 - Anterior telopod with 4 telopoditomeres.................................................................... 5 2. Tergites shiny, glabrous. Process of podomere 2 of posterior telopod swollen only at inner margin. Endotergum with 3–4 dense rows of short marginal bristles.................................................... S. clavigerum Verhoeff, 1924 - Tergites hairy, process of podomere 2 of posterior telopod strongly swollen apically, spherical, broader than base of podomere.................................................................................................... 3 3. Inner margin of femur extended, with teeth. Lateral margin of prefemur with a sharp projection juxtaposed to coxal process. Body coloration black. Head smooth. Process of podomere 2 of posterior telopods truncated. Operculum of vulva two-horned, both horns blunt.................................................................... S. bicorne Attems, 1938 - Inner margin of femur smooth, prefemur without a process. Body coloration dirty olive-brown; head punctured and densely hirsute......................................................................... S. hirsutum Verhoeff, 1924 4. Inner margin of femur extended, with teeth ( Figs 2D , 5D ). Locking carina on anal shield short......................... 6 - Inner margin of femur smooth. Locking carina on anal shield very long........................................... 7 5. Body coloration dark olive, posterior margin uniformly light brown, antennae and legs conspicuously orange ( Fig. 1A ). Prefemur without lateral projection ( Fig. 2D ). Apical margins of vulval operculum equally converging towards a mesal tip ( Fig. 2I ). Tip of laterotergite 1 slightly extended, well-rounded. Male antennomere 6 thickened ( Fig. 3A )......... S. cattiense sp. nov. - Body coloration dark chestnut brown, posterior tergal margins black, legs dark brown, tips light brown ( Fig. 1B ). Lateral margin of prefemur with a sharp projection juxtaposed to coxal process ( Fig. 5D ). Apical margin of vulval operculum recessed centrally ( Fig. 5H ). Tip of laterotergite 1 slightly extended, sharp. Male antennomere 6 axe-shaped ( Fig. 6A ).............................................................................................. S. konkakinhense sp. nov. 6. Podomere 3 of anterior telopod distally with a lateral extension; podomere 4 of posterior telopod distally with an incurved process overlapping with process of podomere 2. Marginal setae of terga not reaching posterior margin.................................................................................................... S. separatum Attems, 1953 - Podomere 3 of anterior telopod cylindrical, without lateral extension; podomere 4 of posterior telopod without incurved pro- cess, but with 3 separate spines. Marginal setae of terga reaching the posterior margin................................................................................................... S. truncatum Wongthamwanich et al ., 2012