A revision of Biting Midges of the Subgenera Forcipomyia (Metaforcipomyia) and F. (Saliohelea) from Costa Rica (Diptera: Ceratopogonidae) Author Spinelli, Gustavo R. Author Marino, Pablo I. Author Borkent, Art text Zootaxa 2012 3419 1 52 journal article 10.5281/zenodo.215031 215f451e-15f2-440d-b4f1-a5ca804ec0cb 1175-5326 215031 Forcipomyia ( Metaforcipomyia ) atenasensis n. sp. ( Figs. 34–39 , 140 ) Diagnosis. Male: only species of Neotropical Forcipomyia ( Metaforcipomyia ) with a hind tarsal ratio of 0.93, labrum tapering apically, legs darkly pigmented except with the base of the hind tibia paler and parameres absent. Female: unknown. Description. Male. Head. Pale brown. Eyes abutting medially for length of three ommatidia. Antenna ( Fig. 34 ) pale brown with plume setae poorly developed, flagellomeres 2–4 spherical, flagellomere 10 1.5 X longer than flagellomere 11, flagellomeres 10–13 elongate; flagellomere 13 with long apical nipple, not constricted basally; AR 1.15. Palpus ( Fig. 35 ) pale brown; segment 3 swollen proximally, with scattered sensillae on surface; PR 3.06. Labrum tapering apically. Thorax. Pale except scutellum and postscutellum brown; scutellum 8–9 strong setae and few minor ones. Legs ( Fig. 36 ) brown; femora brown with at least ¼ apical pale, tibiae with broad basal portion paler than apex (more evident on hind leg); apex of hind tibia with 5 spines; tarsomeres brown with pectinate scales; tarsomere 1 of foreleg with row of 7, 8 spine-like bristles, tarsomere 1 of hind leg with two rows of thick setae on basal portion; foreleg TR 1.75, midleg TR 0.71, hind leg TR 0.93; claws curved, moderately stout. Wing ( Fig. 37 ) plain, without pattern of pigmented membrane; first radial cell obliterated; second radial cell well developed, fork of cubitus situated at level of apex of costa; wing length 1.76 mm ; breadth 0.60 mm ; CR 0.39. Halter whitish. Abdomen. Tergites pale brown with brownish lateral spots. Genitalia ( Fig. 38 ): Brown. Tergite 9 short, only extending to 1/4 length of gonocoxite; posterior margin concave; cercus lobe-like, produced beyond 1/2 length of gonocoxite; sternite 9 broad, posterior margin nearly straight. Gonocoxite stout, 1.8 X longer than greatest breadth; gonostylus slightly paler, 0.85 length of gonocoxite, almost straight, tip blunt. Parameres absent, gonocoxal apodemes slightly curved. Aedeagus ( Fig. 39 ) stout, abruptly narrowed to elongate, slender directed prong, tip pointed; basal arch poorly sclerotized, extending to 1/10 of total length; lateral arms extending laterally. Female. Unknown. Distribution. This species is known only from the type locality in Costa Rica ( Fig. 140 ). The holotype was swept from near a watering trough, fed by a spring in Atenas, Costa Rica (approximately 200 m NW of Don Yayo, a local grocery store), at an altitude of about 650 meters. Taxonomic discussion. This species is similar to F . anniae in general body pigmentation and aedeagus shape. However, the latter species is easily distinguished by the coloration of the hind leg, with a broad basal dark band on the hind femur and subbasal and apical dark bands on the hind tibia. FIGURES 34–39. Forcipomyia ( Metaforcipomyia ) atenasensis sp. n. , male. 34—flagellum; 35-palpus; 36-legs (lateral view); 37—wing; 38—genitalia (ventral view); 39—aedeagus. Type material. Holotype male on microscope slide, labeled “ HOLOTYPE Forcipomyia (Metaforcipomyia) atenasensis Spinelli, Marino and Borkent , Costa Rica , Atenas, 29-XI-1993 , A. Borkent, CD 1688” ( CNCI ). Derivation of specific epithet. The name atenasensis refers to the type-locality.