Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae) Author Orozco, Jesús Molecular Ecology and Systematics Group Department of Botany and Department of Zoology and Entomology Rhodes University Grahamstown, SOUTH AFRICA 6140 text The Coleopterists Bulletin 2012 mo 11 2012-12-20 66 1 182 journal article 10.1649/0010-066X-66.mo4.1 1938-4394 4907222 Euphoria devulsa Horn, 1880 (Appendix 4: Fig. 5 ) Euphoria devulsa Horn 1880: 402 . Original combination. Lectotype at MCZ designated by Hardy (2001) , examined. Other syntypes not found; no paralectotypes designated. Description ( n = 23). Length 9.3–10.7 mm ; width 5.3–7.3 mm . Color : Body shiny black, elytra occasionally reddish brown, frequently with small, whitish, cretaceous markings. Head : Frons flat, strongly strigose, glabrous to moderately densely setose; setae long, yellowish. Clypeus short, subquadrate, strongly strigopunctate, apex and sides not raised, apex vaguely sinuate in dorsal view. Antennal club shorter than stem, subequal in length in both sexes. Pronotum : Surface densely punctate; punctures lunulate, small to moderate in size, denser towards sides, glabrous to moderately densely setose; setae short to long, yellowish. Base in front of scutellum vaguely to weakly emarginate. Scutellum longer than wide, densely punctate; punctures lunulate, setose. Elytra : Surface densely punctate; striae bearing 3 evenly spaced rows of lunulate punctures, grooves occasionally present. Costae weakly to moderately raised. Posterior half of sutural costa raised in lateral view. Pygidium : Surface concentrically striate, striae discontinuous, glabrous to moderately densely setose; setae short, yellowish. Legs : Apical and medial protibial teeth closer to each other than to basal tooth, females frequently with worn teeth. Mesotibial carina well developed, frequently with 2 sharp, short teeth. Metatibiae moderately expanded apically, metatibial spurs subequal in shape in males, internal spur frequently lanceolate in females. Venter : Mesometasternal process small, extending anteriorly to same level as mesocoxae or slightly beyond, apex variably truncate, moderately densely setose. Mesepimera, metasternum, and metacoxae setose, setae as on legs. Metasternum rugose, setose laterally, glabrous, impunctate at middle. Abdominal sternites laterally setose, abdomen of males flat, slightly convex in females. Male genitalia : Parameres as in Fig. 5c . Diagnosis. Euphoria devulsa is distinguished from other species in the group by the subquadrate clypeus, striae bearing three evenly spaced rows of lunulate punctures, and the apex of the pygidium sexually dimorphic (round in males, acuminate in females). This species is close to E. levinotata but can be separated based on the striae bearing 3–4 irregular rows of grooves, double grooves, lunulate punctures, and geminate punctures, and the pygidium at its apex being round in both sexes in E . levinotata . Taxonomic History. Horn (1880) described E . devulsa based on 40 males and 48 females from “near San Antonio, Texas ”, USA . Casey (1915) included E . devulsa in the subgenus Isorhipina with certain doubts about this placement: “ devulsa Horn is probably not closely related to dimidiata and possibly may not belong to this group at all, but not having seen the species I am unable to do more than give the characters assigned by its author.” Hardy (2001) included the species in his “histrionica group” of Euphoria together with E . histrionica , E . subtomentosa , E . lineoligera , E . biguttata , E . canescens , and E . dimidiata . Natural History. Adults have been collected in flight intercept traps and in nests of N. micropus . Temporal Distribution. February (1), April (1), May (5), June (1), July (2), August (1), September (3), December (2) ( Fig. 5d ). Geographic Distribution. Known from Texas and New Mexico in the United States ( Fig. 5e ) . Specimens Examined (23). Ty p e m a t e r i a l: Euphoria devulsa Horn, 1880 ; lectotype at MCZ labeled “Tex// E ./ devulsa/ Horn// MCZ TYPE/ 1902// TYPE No. 3709/ Euphoria / devulsa/ G . H . Horn// LECTOTYPE / Euphoria / devulsa Horn / 1880/ By A . R . Hardy/ 1977”. Other material: USA (22): NEW MEXICO . Guadalupe Co .: Newkirk (1). TEXAS . Bexar Co .: San Antonio (2); Cameron Co .: Boca Chica (1), Brownsville (3); Dallas Co .: No data (1); Kenedy Co .: Laguna Madre (2); Kerr Co .: Kerrville (1); Menard Co .: Menard (1); Nueces Co .: Corpus Christi (9); Travis Co .: Austin (1).