<document id="3EE544FE6A1C4D13100C5E56DD5617AA" ID-DOI="10.1080/00222933.2018.1536813" ID-ISSN="1464-5262" ID-Zenodo-Dep="5177445" ID-ZooBank="A0A6251E-DFD0-4BB7-90EE-D7FA29AE8243" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628617027899" checkinUser="felipe" docAuthor="Mattos, Ingrid &amp; Miras Mermudes, José Ricardo" docDate="2018" docId="76348783FFE8FFCDDF0AFD00AB3D04A7" docLanguage="en" docName="JNatHist.52.37-38.2351-2367.pdf" docOrigin="Journal of Natural History 52 (37 - 38)" docSource="http://dx.doi.org/10.1080/00222933.2018.1536813" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Passalus (Passalus) aculeatus Percheron 1835" docType="treatment" docVersion="4" lastPageNumber="2363" masterDocId="8A0DFFFBFFE5FFC0DE7AFFE5A875005E" masterDocTitle="A new species of Passalus (Passalus) from Atlantic Rainforest, with a key and checklist for the Brazilian Petrejus group (Coleoptera, Passalidae)" masterLastPageNumber="2367" masterPageNumber="2351" pageNumber="2363" updateTime="1699047652107" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FC7F9FBF284EA1DD52E91143B909314B">A new species of Passalus (Passalus) from Atlantic Rainforest, with a key and checklist for the Brazilian Petrejus group (Coleoptera, Passalidae)</mods:title>
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<mods:namePart id="00ADA96FA8F6F88957BA1F7702316346">Mattos, Ingrid</mods:namePart>
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<mods:namePart id="DBF95178B125C25BBB1BEE717B79701D">Miras Mermudes, José Ricardo</mods:namePart>
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<mods:title id="EAAD30D5E0AE7CD1AF5C88C8BA91B310">Journal of Natural History</mods:title>
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<mods:date id="77FAB6EB6E7ADA5A823EAFE5506AC440">2018</mods:date>
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<mods:number id="32774A12B3B315897E74C0C60B8DEA08">2018-11-06</mods:number>
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<mods:number id="DF577DA261CE3ACB4BA227208F78E4F6">52</mods:number>
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<mods:number id="3D32AD4B23874315076BD28805660990">37 - 38</mods:number>
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<treatment id="76348783FFE8FFCDDF0AFD00AB3D04A7" ID-DOI="http://doi.org/10.5281/zenodo.5188731" ID-Zenodo-Dep="5188731" LSID="urn:lsid:plazi:treatment:76348783FFE8FFCDDF0AFD00AB3D04A7" httpUri="http://treatment.plazi.org/id/76348783FFE8FFCDDF0AFD00AB3D04A7" lastPageNumber="2363" pageId="13" pageNumber="2363">
<subSubSection id="B687651EFFE8FFCDDF0AFD00AA1C031F" pageId="13" pageNumber="2363" type="nomenclature">
<paragraph id="FE223695FFE8FFCDDF0AFD00ABC602A0" blockId="13.[368,947,741,766]" box="[368,947,741,766]" pageId="13" pageNumber="2363">
<heading id="A56A81F9FFE8FFCDDF0AFD00ABC602A0" bold="true" box="[368,947,741,766]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="2363" reason="3">
<taxonomicName id="399D4D16FFE8FFCDDF0AFD00ABC602A0" ID-CoL="4DXNF" authority="Percheron, 1835" authorityName="Percheron" authorityYear="1835" box="[368,947,741,766]" class="Insecta" family="Passalidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="2363" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Passalus">
<emphasis id="CCE9EA87FFE8FFCDDF0AFD00ABC602A0" bold="true" box="[368,947,741,766]" pageId="13" pageNumber="2363">
<emphasis id="CCE9EA87FFE8FFCDDF0AFD00A9AC02A0" bold="true" box="[368,473,741,766]" italics="true" pageId="13" pageNumber="2363">Passalus</emphasis>
(
<emphasis id="CCE9EA87FFE8FFCDDF91FD00AA2102A0" bold="true" box="[491,596,741,766]" italics="true" pageId="13" pageNumber="2363">Passalus</emphasis>
) 
<emphasis id="CCE9EA87FFE8FFCDDC1FFD00AAAF02A0" bold="true" box="[613,730,741,766]" italics="true" pageId="13" pageNumber="2363">aculeatus</emphasis>
Percheron, 1835
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="FE223695FFE8FFCDDEDAFCCCAA1C031F" blockId="13.[160,617,809,833]" box="[160,617,809,833]" pageId="13" pageNumber="2363">
Type-locality: 
<collectingCountry id="868A7605FFE8FFCDDF47FCCCA90C031F" box="[317,377,809,833]" name="Brazil" pageId="13" pageNumber="2363">Brazil</collectingCountry>
(Collection M. Gory).
</paragraph>
</subSubSection>
<subSubSection id="B687651EFFE8FFCDDEDAFC9CAC7003EB" box="[160,1029,889,949]" pageId="13" pageNumber="2363" type="distribution">
<paragraph id="FE223695FFE8FFCDDEDAFC9CA95903CC" blockId="13.[160,1029,889,949]" box="[160,300,889,914]" pageId="13" pageNumber="2363">
<emphasis id="CCE9EA87FFE8FFCDDEDAFC9CA95903CC" bold="true" box="[160,300,889,914]" italics="true" pageId="13" pageNumber="2363">Distribution</emphasis>
</paragraph>
<paragraph id="FE223695FFE8FFCDDEDAFC78AC7003EB" blockId="13.[160,1029,889,949]" box="[160,1029,925,949]" pageId="13" pageNumber="2363">
<materialsCitation id="4EF53CC8FFE8FFCDDEDAFC78AC7403EB" box="[160,1025,925,949]" country="Brazil" location="Rio de Janeiro" pageId="13" pageNumber="2363" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingCountry id="868A7605FFE8FFCDDEDAFC78A8A903EB" box="[160,220,925,949]" name="Brazil" pageId="13" pageNumber="2363">Brazil</collectingCountry>
(
<collectingRegion id="3C59F877FFE8FFCDDE91FC78A9E403EB" box="[235,401,925,949]" country="Brazil" name="Rio de Janeiro" pageId="13" pageNumber="2363">Rio de Janeiro</collectingRegion>
, 
<collectingRegion id="3C59F877FFE8FFCDDFE6FC78AA7A03EB" box="[412,527,925,949]" country="Brazil" name="Sao Paulo" pageId="13" pageNumber="2363">São Paulo</collectingRegion>
, 
<collectingRegion id="3C59F877FFE8FFCDDC61FC78AA1F03EB" box="[539,618,925,949]" country="Brazil" name="Parana" pageId="13" pageNumber="2363">Paraná</collectingRegion>
, 
<collectingRegion id="3C59F877FFE8FFCDDC0FFC78AB6903EB" box="[629,796,925,949]" country="Brazil" name="Santa Catarina" pageId="13" pageNumber="2363">Santa Catarina</collectingRegion>
, 
<collectingRegion id="3C59F877FFE8FFCDDD52FC78AB8E03EB" box="[808,1019,925,949]" country="Brazil" name="Rio Grande do Sul" pageId="13" pageNumber="2363">Rio Grande do Sul</collectingRegion>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B687651EFFE8FFCDDEDAFC08AB3D04A7" pageId="13" pageNumber="2363" type="materials_examined">
<paragraph id="FE223695FFE8FFCDDEDAFC08A9F50458" blockId="13.[160,1157,1005,1273]" box="[160,384,1005,1030]" pageId="13" pageNumber="2363">
<emphasis id="CCE9EA87FFE8FFCDDEDAFC08A9F50458" bold="true" box="[160,384,1005,1030]" italics="true" pageId="13" pageNumber="2363">Material examined</emphasis>
</paragraph>
<paragraph id="FE223695FFE8FFCDDEDAFBF4AB3D04A7" blockId="13.[160,1157,1005,1273]" pageId="13" pageNumber="2363">
<materialsCitation id="4EF53CC8FFE8FFCDDEDAFBF4AC510415" collectingDate="2012-03-23" collectionCode="DZRJ" collectorName="Sino" country="Brazil" county="Teresopolis" elevation="1265" latitude="-22.449389" location="Orgaos" longLatPrecision="1" longitude="-43.00397" municipality="Parque Nacional da Serra" pageId="13" pageNumber="2363" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingCountry id="868A7605FFE8FFCDDEDAFBF4A8870477" box="[160,242,1041,1065]" name="Brazil" pageId="13" pageNumber="2363">BRAZIL</collectingCountry>
. 
<collectingRegion id="3C59F877FFE8FFCDDF7AFBF4A9DC0477" box="[256,425,1041,1065]" country="Brazil" name="Rio de Janeiro" pageId="13" pageNumber="2363">Rio de Janeiro</collectingRegion>
: 
<collectingCounty id="17434E19FFE8FFCDDFCDFBF4AA430477" box="[439,566,1041,1065]" pageId="13" pageNumber="2363">Teresópolis</collectingCounty>
(
<collectingMunicipality id="1E46ACEFFFE8FFCDDC3DFBF4AB050477" box="[583,880,1041,1065]" pageId="13" pageNumber="2363">Parque Nacional da Serra</collectingMunicipality>
dos 
<location id="FB42604EFFE8FFCDDDD4FBF4AC740477" LSID="urn:lsid:plazi:treatment:76348783FFE8FFCDDF0AFD00AB3D04A7:FB42604EFFE8FFCDDDD4FBF4AC740477" box="[942,1025,1041,1065]" country="Brazil" county="Teresopolis" latitude="-22.449389" longLatPrecision="1" longitude="-43.00397" municipality="Parque Nacional da Serra" name="Orgaos" pageId="13" pageNumber="2363" stateProvince="Rio de Janeiro">Órgãos</location>
, PARNASO, trilha do 
<collectorName id="53685343FFE8FFCDDF77FBD6A9350415" box="[269,320,1075,1099]" pageId="13" pageNumber="2363">Sino</collectorName>
, 
<quantity id="39659B70FFE8FFCDDF34FBD6A9D20415" box="[334,423,1075,1099]" metricMagnitude="3" metricUnit="m" metricValue="1.265" pageId="13" pageNumber="2363" unit="m" value="1265.0">
<elevation id="75B0D1A6FFE8FFCDDF34FBD6A9D20415" box="[334,423,1075,1099]" metricMagnitude="3" metricUnit="m" metricValue="1.265" pageId="13" pageNumber="2363" unit="m" value="1265.0">1265 m</elevation>
</quantity>
, 
<geoCoordinate id="9BA95052FFE8FFCDDFCFFBD6AA310415" box="[437,580,1075,1099]" degrees="22" direction="south" minutes="26" orientation="latitude" pageId="13" pageNumber="2363" precision="1" seconds="57.8" value="-22.449389">
22°26 
<emphasis id="CCE9EA87FFE8FFCDDF8FFBD6A98E0415" bold="true" box="[501,507,1075,1099]" italics="true" pageId="13" pageNumber="2363">ʹ</emphasis>
57.8 
<emphasis id="CCE9EA87FFE8FFCDDC53FBD6AA410415" bold="true" box="[553,564,1075,1099]" italics="true" pageId="13" pageNumber="2363">″</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="9BA95052FFE8FFCDDC2BFBD6AA9F0415" box="[593,746,1075,1099]" degrees="43" direction="west" minutes="00" orientation="longitude" pageId="13" pageNumber="2363" precision="1" seconds="14.3" value="-43.00397">
43°00 
<emphasis id="CCE9EA87FFE8FFCDDCEBFBD6AAE20415" bold="true" box="[657,663,1075,1099]" italics="true" pageId="13" pageNumber="2363">ʹ</emphasis>
14.3 
<emphasis id="CCE9EA87FFE8FFCDDCBFFBD6AAA50415" bold="true" box="[709,720,1075,1099]" italics="true" pageId="13" pageNumber="2363">″</emphasis>
W
</geoCoordinate>
), 2, 
<date id="8A231055FFE8FFCDDD66FBD6ABB90415" box="[796,972,1075,1099]" pageId="13" pageNumber="2363" value="2012-03-23">
<collectingDate id="9A67E9BDFFE8FFCDDD66FBD6ABB90415" box="[796,972,1075,1099]" pageId="13" pageNumber="2363" value="2012-03-23">23 March 2012</collectingDate>
</date>
, (
<collectionCode id="988CAE50FFE8FFCDDDA5FBD6AC6B0415" box="[991,1054,1075,1099]" pageId="13" pageNumber="2363">DZRJ</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="4EF53CC8FFE8FFCDDA4BFBD6A9CF04CF" collectingDate="1937-12" collectingDateMax="1937-12-04" collectingDateMin="1937-12" collectorName="Coll Claretiano" country="Brazil" location="Curitiba" pageId="13" pageNumber="2363" specimenCount="1" stateProvince="Parana">
<collectingRegion id="3C59F877FFE8FFCDDA4BFBD6ACF50415" box="[1073,1152,1075,1099]" country="Brazil" name="Parana" pageId="13" pageNumber="2363">Paraná</collectingRegion>
: 
<location id="FB42604EFFE8FFCDDEDAFBB3A88E0430" LSID="urn:lsid:plazi:treatment:76348783FFE8FFCDDF0AFD00AB3D04A7:FB42604EFFE8FFCDDEDAFBB3A88E0430" box="[160,251,1110,1134]" country="Brazil" name="Curitiba" pageId="13" pageNumber="2363" stateProvince="Parana">Curitiba</location>
, 5, 
<date id="8A231055FFE8FFCDDF5EFBB3A9AF0430" box="[292,474,1110,1134]" pageId="13" pageNumber="2363" value="1937-12">
<collectingDate id="9A67E9BDFFE8FFCDDF5EFBB3A9AF0430" box="[292,474,1110,1134]" pageId="13" pageNumber="2363" value="1937-12">December 1937</collectingDate>
</date>
, 5, 
<date id="8A231055FFE8FFCDDC79FBB3AAA40430" box="[515,721,1110,1134]" pageId="13" pageNumber="2363" value="1937-12-03">
<collectingDate id="9A67E9BDFFE8FFCDDC79FBB3AAA40430" box="[515,721,1110,1134]" pageId="13" pageNumber="2363" value="1937-12-03">3 December 1937</collectingDate>
</date>
, 1, 
<date id="8A231055FFE8FFCDDC80FBB3ABB20430" box="[762,967,1110,1134]" pageId="13" pageNumber="2363" value="1937-12-04">
<collectingDate id="9A67E9BDFFE8FFCDDC80FBB3ABB20430" box="[762,967,1110,1134]" pageId="13" pageNumber="2363" value="1937-12-04">4 December 1937</collectingDate>
</date>
, 
<collectorName id="53685343FFE8FFCDDDAEFBB3AC0E0430" box="[980,1147,1110,1134]" pageId="13" pageNumber="2363">Coll Claretiano</collectorName>
/ 
<taxonomicName id="399D4D16FFE8FFCDDEDAFB9DA9CF04CF" authority="Perch. Pereira" authorityName="Perch. Pereira" box="[160,442,1144,1169]" class="Insecta" family="Passalidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="2363" phylum="Arthropoda" rank="species" species="aculeatus">
<emphasis id="CCE9EA87FFE8FFCDDEDAFB9DA91B04CF" box="[160,366,1144,1169]" italics="true" pageId="13" pageNumber="2363">Passalus aculeatus</emphasis>
Perch
</taxonomicName>
</materialsCitation>
. Pereira det. 
<materialsCitation id="4EF53CC8FFE8FFCDDC29FB9CA9CD04ED" collectingDate="1937-12-03" collectionCode="IEXA" collectorName="Coll Claretiano" country="Brazil" pageId="13" pageNumber="2363" specimenCount="1">
940 (
<collectionCode id="988CAE50FFE8FFCDDCF6FB9CAAB004CF" box="[652,709,1145,1169]" pageId="13" pageNumber="2363">IEXA</collectionCode>
), 1, 
<date id="8A231055FFE8FFCDDC8CFB9CABB004CF" box="[758,965,1145,1169]" pageId="13" pageNumber="2363" value="1937-12-03">
<collectingDate id="9A67E9BDFFE8FFCDDC8CFB9CABB004CF" box="[758,965,1145,1169]" pageId="13" pageNumber="2363" value="1937-12-03">3 December 1937</collectingDate>
</date>
, 
<collectorName id="53685343FFE8FFCDDDA9FB9CAC0E04CF" box="[979,1147,1145,1169]" pageId="13" pageNumber="2363">Coll Claretiano</collectorName>
/ 
<taxonomicName id="399D4D16FFE8FFCDDEDAFB7FA9CD04ED" authority="Perch. Pereira" authorityName="Perch. Pereira" box="[160,440,1178,1203]" class="Insecta" family="Passalidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="2363" phylum="Arthropoda" rank="species" species="aculeatus">
<emphasis id="CCE9EA87FFE8FFCDDEDAFB7FA91804ED" box="[160,365,1178,1203]" italics="true" pageId="13" pageNumber="2363">Passalus aculeatus</emphasis>
Perch
</taxonomicName>
</materialsCitation>
. Pereira det. 939 (
<collectionCode id="988CAE50FFE8FFCDDCFCFB7EAAC904ED" box="[646,700,1179,1203]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="13" pageNumber="2363">MLP</collectionCode>
). 
<materialsCitation id="4EF53CC8FFE8FFCDDCB4FB7EA9030488" collectingDate="1947-04" collectingDateMax="1947-12-01" collectingDateMin="1947-04" country="Brazil" location="Pinhal" pageId="13" pageNumber="2363" specimenCount="1" stateProvince="Santa Catarina">
<collectingRegion id="3C59F877FFE8FFCDDCB4FB7EAB0004ED" box="[718,885,1179,1203]" country="Brazil" name="Santa Catarina" pageId="13" pageNumber="2363">Santa Catarina</collectingRegion>
: 
<location id="FB42604EFFE8FFCDDDF8FB7EABBF04ED" LSID="urn:lsid:plazi:treatment:76348783FFE8FFCDDF0AFD00AB3D04A7:FB42604EFFE8FFCDDDF8FB7EABBF04ED" box="[898,970,1179,1203]" country="Brazil" name="Pinhal" pageId="13" pageNumber="2363" stateProvince="Santa Catarina">Pinhal</location>
, 2, 
<date id="8A231055FFE8FFCDDD88FB7EAC1C04ED" box="[1010,1129,1179,1203]" pageId="13" pageNumber="2363" value="1947-04">
<collectingDate id="9A67E9BDFFE8FFCDDD88FB7EAC1C04ED" box="[1010,1129,1179,1203]" pageId="13" pageNumber="2363" value="1947-04">April 1947</collectingDate>
</date>
; 
<date id="8A231055FFE8FFCDDA0FFB7EA9230488" pageId="13" pageNumber="2363" value="1947-12-01">
<collectingDate id="9A67E9BDFFE8FFCDDA0FFB7EA9230488" pageId="13" pageNumber="2363" value="1947-12-01">1 December 1947</collectingDate>
</date>
, A
</materialsCitation>
. 
<materialsCitation id="4EF53CC8FFE8FFCDDFF9FB5BAA680488" box="[387,541,1214,1238]" country="Brazil" location="Maller" pageId="13" pageNumber="2363" specimenCount="1">
<location id="FB42604EFFE8FFCDDFF9FB5BA9BD0488" LSID="urn:lsid:plazi:treatment:76348783FFE8FFCDDF0AFD00AB3D04A7:FB42604EFFE8FFCDDFF9FB5BA9BD0488" box="[387,456,1214,1238]" country="Brazil" name="Maller" pageId="13" pageNumber="2363">Maller</location>
ex-coll
</materialsCitation>
. 
<materialsCitation id="4EF53CC8FFE8FFCDDC53FB5BAB1B0488" box="[553,878,1214,1238]" collectionCode="IEXA" country="Brazil" location="Frank Johnson Donor" pageId="13" pageNumber="2363" specimenCount="1">
<location id="FB42604EFFE8FFCDDC53FB5BAB6A0488" LSID="urn:lsid:plazi:treatment:76348783FFE8FFCDDF0AFD00AB3D04A7:FB42604EFFE8FFCDDC53FB5BAB6A0488" box="[553,799,1214,1238]" country="Brazil" name="Frank Johnson Donor" pageId="13" pageNumber="2363">Frank Johnson Donor</location>
(
<collectionCode id="988CAE50FFE8FFCDDD54FB5BAB120488" box="[814,871,1214,1238]" pageId="13" pageNumber="2363">IEXA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="4EF53CC8FFE8FFCDDD00FB5BAA1504A7" collectingDate="1937-01" country="Brazil" location="Sao Francisco de Paula" pageId="13" pageNumber="2363" specimenCount="1" stateProvince="Rio Grande do Sul">
<collectingRegion id="3C59F877FFE8FFCDDD00FB5BAC3B0488" box="[890,1102,1214,1238]" country="Brazil" name="Rio Grande do Sul" pageId="13" pageNumber="2363">Rio Grande do Sul</collectingRegion>
: 
<location id="FB42604EFFE8FFCDDA20FB5BA90204A7" LSID="urn:lsid:plazi:treatment:76348783FFE8FFCDDF0AFD00AB3D04A7:FB42604EFFE8FFCDDA20FB5BA90204A7" country="Brazil" name="Sao Francisco de Paula" pageId="13" pageNumber="2363" stateProvince="Rio Grande do Sul">São Francisco de Paula</location>
, 22, 
<date id="8A231055FFE8FFCDDFD6FB04AA3104A7" box="[428,580,1249,1273]" pageId="13" pageNumber="2363" value="1937-01">
<collectingDate id="9A67E9BDFFE8FFCDDFD6FB04AA3104A7" box="[428,580,1249,1273]" pageId="13" pageNumber="2363" value="1937-01">January 1937</collectingDate>
</date>
, P
</materialsCitation>
<materialsCitation id="4EF53CC8FFE8FFCDDC1AFB04AA0604A7" box="[608,627,1249,1273]" collectingDate="1937-01" country="Brazil" isTailEnumeration="true" location="Sao Francisco de Paula" pageId="13" pageNumber="2363" specimenCount="1" stateProvince="Rio Grande do Sul">.P</materialsCitation>
. Buck col. (
<collectionCode id="988CAE50FFE8FFCDDC89FB04AB4A04A7" box="[755,831,1249,1273]" pageId="13" pageNumber="2363">CEIOC</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>