Review of Dissomphalus Ashmead, 1893 (Hymenoptera, Bethylidae) from Espíri- to Santo, Brazil, with description of twenty-one new species Author Colombo, Wesley D. Author Azevedo, Celso O. text Zootaxa 2016 4143 1 1 84 journal article 10.11646/zootaxa.4143.1.1 1b4c3fd4-ecd4-4d0f-9bd4-a300eacd5f27 1175-5326 260759 0DD2D30A-999D-4DFE-90F8-0E809733C70F secretus species-group, new species group Diagnosis. Male. T2 with lateral depressions or not, well-defined side pits, circular-shaped, higher edges, tubercle below surface of tergite, many setae at apex, bristles on edge depressions or not. Remarks. This species-group contains four species: Dissomphalus cacirus sp. nov. , Dissomphalus mirim sp. nov. and Dissomphalus secretus sp. nov. , all from Espírito Santo . Distribution. Brazil ( Espírito Santo ).