Redescriptions Of Three Species Of The Genus Neobisium From The Balkans And The Carpathians (Arachnida: Pseudoscorpiones) Author Novák, János Author Hörweg, Christoph text Acta Zoologica Academiae Scientiarum Hungaricae 2017 2017-07-21 63 3 277 292 http://dx.doi.org/10.17109/azh.63.3.277.2017 journal article 10.17109/AZH.63.3.277.2017 2064-2474 39196813-82D8-4028-98A1-D927D3D60E65 Neobisium (Neobisium) bosnicum Beier, 1939 (Figs 4A–F) Typematerialexamined ( syntypes ): 1 m , 2 f, BosniaandHerzegovina, Bjelašnica Planina, cleavecave, southeasttoSarajevo, 2067 masl ., 29.06.1912 (NHMW-25213); 3 m , 2 f, BosniaandHerzegovina, fromcavesoftheBjelašnicaMts, 1918 (NHMW-25214); 1 m , 1 f, BosniaandHerzegovina, VranPlanina, westtoNarenta, understones, 1922 (NHMW- 25215). Description. Medium-sizedpseudoscorpionwithepigeanhabitus. Carapaceandter- gitesolivebrown, pedipalpsreddish- brown, otherpartsofopisthosomaandlegsyellow- ish. MeasurementsandratiosofthestudiedspecimensasinTable 3. Carapacelongerthanbroad, surfacesmooth (Fig. 4A). Epistometriangularand prominent. Twopairsofwelldevelopedeyeswithlens. Onepairofslitlikelyrifissuresat anteriorandtwopairsatposteriorhalfofcarapace. Preocularsetaeabsent. Setalformula: 4:6:6–10:8–14 (26–34). Coxalarea. Manducatoryprocesswith 5–6 setae, restofpedipalpalcoxawith 9–14 se- tae; pedalcoxaIwith 10–13 setae; II: 7–14; III: 6–11; IV: 13–17. Anterolateralprocessofcoxa Ilong, triangularandapicallypointed; medialprocessroundandflattenedanddentate. Opisthosomawithgranulostriatepleuralmembrane. ChaetotaxyoftergitesI–X: 8–10:8–16:10–14:12–20:14–20:14–18:12–18:12–16:10–16:10–14. SegmentXIwith 14–22 setae, 6 ofthemtactile. Analconewith 2 ventraland 2 dorsalsetae. Malegenitalregion: sternite II with 6–13 small setae, sternite III with 24–28 setae, of which 12–15 along posterior mar- gin. Inmaleopening 4+4 internalsetae. Medialgenitalsackcouldnotbeclearlyobserved. Female genital region: sternite II with 7–12 small setae, sternite III with 10–14 small setae in a row. Chaetotaxy of sternites IV–X: 15–23:14–26:16–26:14–18:17–22:16–20:17–20. Chelicerawith 6–8 setaeonhand (Fig. 4B), thisnumbercanvarybetweenthecheli- ceraeofthesameexemplar. Subgalealseta 0.59–0.65× frombaseofmovablefinger. Spin- neretaflattenedhyalinetuberclewith 10–14 silkducts. Fixedfingerwith 11–15 smallteeth, movablefingerwith 8–10 smallandmedium-sizedteeth. Rallumwith 8 blades, twodistalmostbladesunilaterallypinnateonanteriorface. Thedistalmostbladewithenlargedbasis andseparatedfromtheproceedingone. Serrulaexteriorwith 24–28 blades, serrulainterior with 22–26 blades. Pedipalp (Fig. 4C) surfacessmooth. Trochanterwithtubercle. Femur 3.58–3.75, patella 2.28–2.50 timesaslongasbroad. Patellawith 3 greaterlyrifissuresonbasalpartand 2 smallernearthearticulationwiththehand. Pedicelofpatellabears 2–4 micropores. Che- la (Fig. 4D). Pedicelbears 3–4 dorsalmicropores. Chelalfingersapproximatelyaslong ashandwithpedicel. Fixedchelalfingerwith 55–63 close- setandpointedteethofequal length (Figs 4E–F). Movablechelalfingerwith 49–54 roundedteeth, dentallineending betweentheleveloftrichobothria sb and b . Trichobothriaofchelalfingersdistributedas inFig. 4D. Distancebetweentrichobothria ib and ist approximatelyaslongasbetween ist and it . Legswithsmoothsurfaces. Clawssmoothandsimple, aroliashorterthanclawson alllegs. SubterminalsetaoftelotarsusIVwithtworami. Tactilesetaepresentontibia, basitarsusandtelotarsusoflegIV. Table 3. Measurements (in mm) and ratios of Neobisium (N.) bosnicum and N. (N.) herzegovinense syntypes.

N. (N.) bosnicum		N. (N.) herzegovinense
Male	Female	Male	Female
Measurements	Ratio	Measurements	Ratio	Measurements	Ratio	Measurements	Ratio
Body	2.69–2.97	2.82–3.01	2.95	3.23–3.35
Carapace	0.62–0.74/0.73–0.89	0.79–0.87	0.68–0.72/0.90–0.95	0.75–0.78	0.73/0.79	0.78–0.79/0.86–0.90
Chelicera
Hand	0.41–0.45/0.24–0.28	0.45–0.51/0.26–0.29	0.45–0.46/0.25–0.26	0.50–0.52/0.28–0.29
Finger	0.27–0.30	0.28–0.34	0.30–0.32	0.33–0.35
Palp
Femur	0.73–0.83/0.20–2.22	3.58–3.75	0.80–0.85/0.22–0.24	3.58–3.72	0.90–0.92/0.20–0.21	4.29–4.60	0.93–1.01/0.22–0.23	4.22–4.24
Patella	0.57–0.66/0.25–0.28	2.28–2.50	0.63–0.68/0.27–0.29	2.23–2.44	0.74–0.75/0.25–0.26	2.88–2.96	0.75–0.77/0.27–0.29	2.65–2.78
Chela	1.29–1.39	1.41–1.46	1.61–1.64	1.65–1.67
Hand	0.51–0.58/0.36–0.40	1.42–1.45	0.58–0.60/0.41–0.42	1.41–1.46	0.65–0.68/0.39	1.67–1.74	0.67–0.72/0.42–0.43	1.59–1.67
Pedicel	0.08–0.10	0.08–0.10	0.13	0.12–0.13
Finger	0.70–0.76	0.75–0.79	0.87	0.84–0.92
Leg I
Trochanter	0.18–0.22/0.14	1.29–1.56	0.21/0.15	1.42	0.23–0.24/0.13	1.78–1.92	0.21–0.24/0.15	1.43–1.64
Femur	0.39–0.41/0.09–0.10	3.92–4.54	0.41–0.42/0.08–0.11	3.87–5.12	0.38–0.40/0.09	4.21–4.45	0.45/0.09–0.10	4.51–5.04
Patella	0.30–0.31/0.10	2.99–3.12	0.30–0.32/0.10	2.97–3.21	0.34–0.36/0.10–0.11	3.28–3.42	0.32–0.35/0.11	2.91–3.20
Tibia	0.27/0.09	3.05	0.29–0.31/0.09–0.10	3.09–3.22	0.31–0.32/0.09	3.43–3.56	0.34–0.35/0.09–0.10	3.76–3.89
Basitarsus	0.21/0.07	2.98	0.20–0.22/0.07–0.09	2.43–2.86	0.20–0.21/0.08	2.51–2.64	0.20–0.21/0.08	2.49–2.63
Telotarsus	0.28/0.06	4.31	0.22–0.24/0.07–0.08	3.02–3.15	0.33–0.34/0.07	4.71–4.84	0.24–0.32/0.06–0.07	4.04–4.58
Leg IV
Trochanter	0.31–0.32/0.16–0.18	1.92–2.02	0.29–0.31/0.17–0.18	1.63–1.83	0.35/0.15	2.33	0.34/0.17	2.11
Femur+patella	0.67–0.72/0.20–0.22	3.36–3.54	0.73–0.75/0.21–0.24	3.12–3.48	0.79/0.20	3.96	0.81/0.20	4.06
Tibia	0.51–0.54/0.12–0.13	4.15–4.26	0.58–0.61/0.14–0.15	3.88–4.34	0.72/0.11	6.55	0.73/0.12	6.08
Basitarsus	0.25–0.29/0.09	2.78–3.22	0.25–0.28/0.10	2.53–2.84	0.31–0.32/0.09	3.43–3.54	0.34/0.10	3.42
Telotarsus	0.32–0.35/0.07–0.08	4.38–4.56	0.33–0.34/0.08	4.14–4.25	0.42/0.09	4.66	0.40/0.09	4.43

Remarks. N. bosnicum wasdescribedbyBeierfrom 4 male , 6 femaleand 1 juvenilesyntypes, fromthreedifferentlocalitiesinBosniaandHerzegovina ( BEIER 1939b ). In 1973 anewsubspecies, N. bosnicumondriasi wasdescribed Fig. 4. Neobisium (N.) bosnicum : A = carapace, male, B = leftchelicera, male, C = rightpedi- palp, female, D = leftchela, male, E = chelalteethoffixedfingeratthelevelof ist , male, F = chelal teeth of fixed finger at the level of et , male; G–J = Neobisium (N.) herzegovinense : G = carapace, female, H = leftchelicera, female, I = rightpedipalp, female, J = rightchela, female fromGreece ( MaHNERT 1973 ). Afterwardsnofurtheroccurrencesofthespe- cieswerepublished. Duringthepresentinvestigationonlytheadultsyntypeswerestudied. Wefoundgreatervariabilityinthesetationofthecarapaceandthetergites, andinthemeasurementdataandproportionsthanpreviouslymentionedin theliterature ( BEIER 1939b , 1963 ). Itisworthtomention, thatinthesample collectedincavesoftheBjelašnicaMtstheoriginaldescriptionreports 2 males and 3 females ; incontrast, wefound 3 malesand 2 femalesinthevial.