<document id="9678EF3A5B284400DE9666076CBA2935" ID-DOI="10.1111/j.1096-3642.2009.00608.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5439924" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630445345639" checkinUser="valdenar" docAuthor="Prendini, Lorenzo &amp; Esposito, Lauren A." docDate="2010" docId="A633A44EFF92A26BFC67A5FBD787B343" docLanguage="en" docName="j.1096-3642.2009.00608.x.pdf" docOrigin="Zoological Journal of the Linnean Society 159 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2009.00608.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Parabuthus glabrimanus Prendini &amp; Esposito 2010, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="682" masterDocId="5A0ADC36FF9BA262FFE8A058D226B42C" masterDocTitle="A reanalysis of Parabuthus (Scorpiones: Buthidae) phylogeny with descriptions of two new Parabuthus species endemic to the Central Namib gravel plains, Namibia" masterLastPageNumber="710" masterPageNumber="673" pageNumber="682" updateTime="1699273108265" updateUser="plazi">
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<mods:title id="8E4EC09E7A0AA5DE31979F2B8E77AC11">A reanalysis of Parabuthus (Scorpiones: Buthidae) phylogeny with descriptions of two new Parabuthus species endemic to the Central Namib gravel plains, Namibia</mods:title>
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<mods:namePart id="9A63F3E91795BF1FC5523C36AFF3D12A">Prendini, Lorenzo</mods:namePart>
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<taxonomicName id="E99A6EDBFF92A26BFC67A5FBD6FBB194" ID-CoL="4CN52" authority="Prendini &amp; Esposito, 2010" authorityName="Prendini &amp; Esposito" authorityYear="2010" box="[911,1245,1443,1464]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="682" phylum="Arthropoda" rank="species" species="glabrimanus" status="sp. nov.">PARABUTHUS GLABRIMANUS</taxonomicName>
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<paragraph id="2E251558FF92A26BFC8AA59BD75FB1F5" blockId="9.[866,1401,1443,1497]" box="[866,1401,1475,1497]" pageId="9" pageNumber="682">
(
<figureCitation id="B6A109DDFF92A26BFC82A59BD1F4B1F4" box="[874,978,1475,1496]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="4.[144,223,1816,1835]" captionStartId-1="5.[164,243,1662,1681]" captionStartId-2="6.[144,223,1135,1154]" captionStartId-3="6.[806,885,1688,1707]" captionTargetBox-0="[152,1420,201,1779]" captionTargetBox-1="[355,1252,195,1626]" captionTargetBox-2="[245,659,199,1102]" captionTargetBox-3="[904,1325,197,1653]" captionTargetId-0="figure-0@4.[144,1428,193,1786]" captionTargetId-1="figure-0@5.[355,1252,195,1629]" captionTargetId-2="figure-168@6.[241,663,195,1105]" captionTargetId-3="figure-189@6.[903,1325,195,1656]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="6" captionText-0="Figure 1. Map showing the known distributions of Parabuthus glabrimanus sp. nov. (triangles), Parabuthus gracilis Lamoral, 1979 (squares), Parabuthus nanus Lamoral, 1979 (stars) and Parabuthus setiventer sp. nov. (circles) in Namibia and South Africa. Contour interval, 500 m. Major sand systems stippled. Magnified inset shows distributions of P. glabrimanus and P. setiventer in finer detail." captionText-1="Figure 2. See Table 2 for details. The optimal tree retrieved by analyses with equal weights and implied weights under k = 3–6 (Table 2). This topology corresponds to the majority rule (&gt; 50%) consensus of most parsimonious trees (MPTs) obtained by seven analyses in which weighting regime was varied; frequency percentages for nodes that collapsed in the strict consensus of the MPTs obtained by the seven analyses are indicated to the right of relevant nodes. Zero-length branches are collapsed. Solid bars indicate uniquely derived apomorphic character states, whereas empty bars indicate parallel derivations of apomorphic states under accelerated transformation (ACCTRAN) optimization. The number above each bar gives the character number, whereas the number below gives the character state. Branch support values are circled at nodes. Refer to Appendix for character descriptions. The East African species are indicated with asterisks. Alternative topologies for nodes A and B shown in Figures 3 and 4 respectively." captionText-2="Figure 3. Alternative topologies for node A (Fig. 2) retrieved by analysis with equal weights. See Table 2 for details." captionText-3="Figure 4. Alternative topologies for node B (Fig. 2) retrieved by analyses with equal weights and implied weights under k = 3–6 (A) and with implied weights under k = 1 and 2 (B, C). See Table 2 for details." figureDoi-0="http://doi.org/10.5281/zenodo.5439926" figureDoi-1="http://doi.org/10.5281/zenodo.5439930" figureDoi-2="http://doi.org/10.5281/zenodo.5439932" figureDoi-3="http://doi.org/10.5281/zenodo.5439934" httpUri-0="https://zenodo.org/record/5439926/files/figure.png" httpUri-1="https://zenodo.org/record/5439930/files/figure.png" httpUri-2="https://zenodo.org/record/5439932/files/figure.png" httpUri-3="https://zenodo.org/record/5439934/files/figure.png" pageId="9" pageNumber="682">FIGS 1–4</figureCitation>
, 
<figureCitation id="B6A109DDFF92A26BFC0AA59BD601B1F4" box="[994,1063,1475,1497]" captionStart="Figure 5" captionStartId="11.[164,243,1789,1808]" captionTargetBox="[164,1444,196,1759]" captionTargetId="figure-1@11.[164,1445,196,1760]" captionTargetPageId="11" captionText="Figure 5. Parabuthus glabrimanus sp. nov. (A, B) and Parabuthus setiventer sp. nov. (C, D), carapace, dorsal aspect. A, holotype ♂ (SMN 2901). B, paratype ♀ (AMNH). C, holotype ♂ (SMN 2918). D, paratype ♀ (AMNH). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5439936" httpUri="https://zenodo.org/record/5439936/files/figure.png" pageId="9" pageNumber="682">5A, B</figureCitation>
, 
<figureCitation id="B6A109DDFF92A26BFBDFA59BD67CB1F5" box="[1079,1114,1475,1497]" captionStart="Figure 6" captionStartId="12.[144,223,1834,1853]" captionTargetBox="[319,1249,195,1805]" captionTargetId="figure-0@12.[319,1249,195,1805]" captionTargetPageId="12" captionText="Figure 6. Parabuthus glabrimanus sp. nov. (A), Parabuthus gracilis Lamoral, 1979 (B), Parabuthus nanus Lamoral, 1979 (C) and Parabuthus setiventer sp. nov. (D), sternites III–VII, ventral aspect. A, ♂ [AMNH (AH 2164)]. B–D, ♂ (AMNH). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5439938" httpUri="https://zenodo.org/record/5439938/files/figure.png" pageId="9" pageNumber="682">6A</figureCitation>
, 
<figureCitation id="B6A109DDFF92A26BFB80A59BD68DB1F4" box="[1128,1195,1475,1496]" captionStart="Figure 7" captionStartId="13.[164,243,1513,1532]" captionTargetBox="[164,1444,194,1482]" captionTargetId="figure-149@13.[164,1444,194,1482]" captionTargetPageId="13" captionText="Figure 7. Parabuthus glabrimanus sp. nov. (A–C) and Parabuthus setiventer sp. nov. (D–F), metasoma and telson. A, D, ventral aspect. B, C, E, F, dorsal aspect. A, B, holotype ♂ (SMN 2901). C, paratype ♀ (AMNH). D, E, holotype ♂ (SMN 2918). F, paratype ♀ (AMNH). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5439940" httpUri="https://zenodo.org/record/5439940/files/figure.png" pageId="9" pageNumber="682">7A–C</figureCitation>
, 
<figureCitation id="B6A109DDFF92A26BFB52A59BD6ECB1F4" box="[1210,1226,1475,1496]" captionStart="Figure 8" captionStartId="14.[144,223,1858,1877]" captionTargetBox="[230,1337,194,1829]" captionTargetId="figure-0@14.[230,1337,194,1829]" captionTargetPageId="14" captionText="Figure 8. Parabuthus glabrimanus sp. nov., habitus. A, B, holotype ♂ (SMN 2918). C, D, paratype ♀ (AMNH). A, C, dorsal aspect. B, D, ventral aspect. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5439942" httpUri="https://zenodo.org/record/5439942/files/figure.png" pageId="9" pageNumber="682">8</figureCitation>
, 
<figureCitation id="B6A109DDFF92A26BFB31A59BD73AB1F4" box="[1241,1308,1475,1496]" captionStart="Figure 9" captionStartId="15.[164,243,1598,1617]" captionTargetBox="[387,1220,194,1568]" captionTargetId="figure-119@15.[387,1220,194,1568]" captionTargetPageId="15" captionText="Figure 9. Parabuthus gracilis Lamoral, 1979 (A) and Parabuthus glabrimanus sp. nov. (B–E), dextral pedipalp chela, illustrating trichobothrial distribution. A, ♂ [AMNH (AH 4397)], external aspect. B, holotype ♂ (SMN 2901), external aspect. C–E, paratype ♀ (AMNH), external (C), ventral (D), and internal (E) aspects. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5439944" httpUri="https://zenodo.org/record/5439944/files/figure.png" pageId="9" pageNumber="682">9B–E</figureCitation>
, 
<figureCitation id="B6A109DDFF92A26BFAC3A59BD757B1F5" box="[1323,1393,1475,1497]" captionStart-0="Figure 10" captionStart-1="Figure 11" captionStart-2="Figure 12" captionStartId-0="16.[144,223,1026,1045]" captionStartId-1="16.[806,885,940,959]" captionStartId-2="16.[806,885,1824,1843]" captionTargetBox-0="[144,761,197,995]" captionTargetBox-1="[806,1422,197,911]" captionTargetBox-2="[806,1423,1165,1794]" captionTargetId-0="figure-208@16.[144,761,196,995]" captionTargetId-1="figure-244@16.[806,1422,196,911]" captionTargetId-2="figure-286@16.[806,1423,1165,1794]" captionTargetPageId-0="16" captionTargetPageId-1="16" captionTargetPageId-2="16" captionText-0="Figure 10. Parabuthus glabrimanus sp. nov., paratype ♀ (AMNH), dextral pedipalp patella and femur, illustrating trichobothrial distribution. A, patella, dorsal aspect. B, patella, external aspect. C, femur, dorsal aspect. Scale bar = 1 mm." captionText-1="Figure 11. Parabuthus glabrimanus sp. nov., paratype ♀ (AMNH), dextral legs I–IV, tibia, basitarsus, and telotarsus, retrolateral aspect, illustrating setae, ungues, and spurs. A, leg I. B, leg II. C, leg III. D, leg IV. Scale bar = 1 mm." captionText-2="Figure 12. Parabuthus glabrimanus sp. nov., paratype ♂ (AMNH), dextral hemispermatophore, ental (A) and ectal (B) aspects. Scale bar = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5439950" figureDoi-1="http://doi.org/10.5281/zenodo.5439946" figureDoi-2="http://doi.org/10.5281/zenodo.5439952" httpUri-0="https://zenodo.org/record/5439950/files/figure.png" httpUri-1="https://zenodo.org/record/5439946/files/figure.png" httpUri-2="https://zenodo.org/record/5439952/files/figure.png" pageId="9" pageNumber="682">10–12</figureCitation>
)
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<location id="2B454383FF92A26BFB54A698D76CB2FA" LSID="urn:lsid:plazi:treatment:A633A44EFF92A26BFC67A5FBD787B343:2B454383FF92A26BFB54A698D76CB2FA" box="[1212,1354,1728,1750]" country="Namibia" county="Swakopmund District" latitude="-23.133333" longLatPrecision="1257" longitude="15.583333" municipality="Namib-Naukluft Park" name="Bloedkoppie" pageId="9" pageNumber="682" stateProvince="Erongo">Bloedkoppie</location>
, 
<quantity id="E962B8BDFF92A26BFAB3A698D787B2F9" box="[1371,1441,1728,1750]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="682" unit="m" value="500.0">
<elevation id="A5B7F26BFF92A26BFAB3A698D787B2F9" box="[1371,1441,1728,1750]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="682" unit="m" value="500.0">500 m</elevation>
</quantity>
east, 
<geoCoordinate id="4BAE739FFF92A26BFC6AA686D632B2D8" box="[898,1044,1757,1780]" degrees="22" direction="south" minutes="50" orientation="latitude" pageId="9" pageNumber="682" precision="1" seconds="36.1" value="-22.84336">22°50′36.1″S</geoCoordinate>
, 
<geoCoordinate id="4BAE739FFF92A26BFBC3A686D6E6B2D8" box="[1067,1216,1757,1780]" degrees="15" direction="east" minutes="23" orientation="longitude" pageId="9" pageNumber="682" precision="1" seconds="22.2" value="15.3895">15°23′22.2″E</geoCoordinate>
, 
<date id="5A243398FF92A26BFB3FA686D7BBB2D8" box="[1239,1437,1758,1780]" pageId="9" pageNumber="682" value="2008-07-31" valueMax="2008-08-02" valueMin="2008-07-31">
<collectingDate id="4A60CA70FF92A26BFB3FA686D7BBB2D8" box="[1239,1437,1758,1780]" pageId="9" pageNumber="682" value="2008-07-31" valueMax="2008-08-02" valueMin="2008-07-31">31.vii–2.viii.2008</collectingDate>
</date>
, 
<collectionCode id="488B8D9DFF92A26BFCD2A6A5D168B33E" box="[826,846,1789,1810]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="682">R</collectionCode>
. 
<collectorName id="836F708EFF92A26BFCBAA6A5D1FCB33F" box="[850,986,1789,1811]" pageId="9" pageNumber="682">A. Wharton</collectorName>
&amp; 
<collectionCode id="488B8D9DFF92A26BFBE9A6A5D635B33E" box="[1025,1043,1789,1810]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="682">T</collectionCode>
. 
<collectorName id="836F708EFF92A26BFBFFA6A5D64AB33F" box="[1047,1132,1789,1811]" pageId="9" pageNumber="682">L. Bird</collectorName>
, 
<quantity id="E962B8BDFF92A26BFB92A6A5D6E7B33E" box="[1146,1217,1789,1810]" metricMagnitude="2" metricUnit="m" metricValue="7.43" pageId="9" pageNumber="682" unit="m" value="743.0">
<elevation id="A5B7F26BFF92A26BFB92A6A5D6E7B33E" box="[1146,1217,1789,1810]" metricMagnitude="2" metricUnit="m" metricValue="7.43" pageId="9" pageNumber="682" unit="m" value="743.0">743 m</elevation>
</quantity>
, under stones, 
<specimenCount id="389CDED1FF92A26BFA9DA6A5D787B33F" box="[1397,1441,1789,1811]" pageId="9" pageNumber="682" type="female">1 ♀</specimenCount>
(SMN 3391); 
<collectorName id="836F708EFF92A26BFC3CA744D685B31D" box="[980,1187,1820,1842]" pageId="9" pageNumber="682">North-east Range</collectorName>
, 
<collectorName id="836F708EFF92A26BFB47A744D6DCB31E" box="[1199,1274,1820,1842]" pageId="9" pageNumber="682">Ganab</collectorName>
flats, 
<geoCoordinate id="4BAE739FFF92A26BFAACA744D7BAB31D" box="[1348,1436,1818,1841]" degrees="23" direction="south" minutes="08" orientation="latitude" pageId="9" pageNumber="682" precision="925" value="-23.133333">23°08′S</geoCoordinate>
, 
<geoCoordinate id="4BAE739FFF92A26BFCD2A762D1B3B37C" box="[826,917,1849,1872]" degrees="15" direction="east" minutes="35" orientation="longitude" pageId="9" pageNumber="682" precision="925" value="15.583333">15°35′E</geoCoordinate>
, 
<date id="5A243398FF92A26BFC43A762D632B37C" box="[939,1044,1850,1872]" pageId="9" pageNumber="682" value="1974-11-08">
<collectingDate id="4A60CA70FF92A26BFC43A762D632B37C" box="[939,1044,1850,1872]" pageId="9" pageNumber="682" value="1974-11-08">8.xi.1974</collectingDate>
</date>
, 
<collectorName id="836F708EFF92A26BFBC2A762D734B37C" box="[1066,1298,1850,1872]" pageId="9" pageNumber="682">S. Endrödy-Younga</collectorName>
, 
<specimenCount id="389CDED1FF92A26BFAC0A762D746B37C" box="[1320,1376,1850,1872]" pageId="9" pageNumber="682" type="male">1 ♂</specimenCount>
[
<specimenCode id="7E3CBD23FF92A26BFA9FA763D15BB342" collectionCode="TM" country="Germany" name="Teylers Museum, Paleontologische" pageId="9" pageNumber="682">TM 11332</specimenCode>
(E-Y 483)]; 
<location id="2B454383FF92A26BFBC9A701D6F5B343" LSID="urn:lsid:plazi:treatment:A633A44EFF92A26BFC67A5FBD787B343:2B454383FF92A26BFBC9A701D6F5B343" box="[1057,1235,1881,1903]" country="Namibia" county="Swakopmund District" latitude="-23.133333" longLatPrecision="1257" longitude="15.583333" municipality="Namib-Naukluft Park" name="Ganab Station" pageId="9" pageNumber="682" stateProvince="Erongo">Ganab Station</location>
, 
<quantity id="E962B8BDFF92A26BFB0FA701D70BB342" box="[1255,1325,1881,1902]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="682" unit="m" value="200.0">
<elevation id="A5B7F26BFF92A26BFB0FA701D70BB342" box="[1255,1325,1881,1902]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="682" unit="m" value="200.0">200 m</elevation>
</quantity>
south of
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>