Review of Glyphidocera Walsingham of Costa Rica (Lepidoptera: Gelechioidea: Glyphidoceridae) Author DAVID ADAMSKI text Zootaxa 2005 858 1 158 journal article 10.5281/zenodo.170793 94a0cedb-15bb-4bfd-955e-77200a536045 1175-5334 170793 511FBF55-10F2-4A00-B15B-CBD34EAB023E Glyphidocera illiterata Meyrick, 1929 (Figs. 150–152, 281, 337 , Map 27 ) Glyphidocera illiterata Meyrick, 1929 : 509 ; Becker, 1984 : 41 (checklist). Diagnosis.— Glyphidocera illiterata can be distinguished from its congeners by a unique combination of features that include a short valva, a stout digitate process of the valva, costal furcae of valva widely separate, and dorsal furca of costa about midway between digitate process of valva and ventral furca. Description.— Head: Vertex dark brown; frontoclypeus dark brown intermixed with few pale­brown scales; outer surface of labial palpus dark brown intermixed with few pale­brown scales to near apical margin of segment II, inner margin dark brown intermixed with few pale­brown scales; scape of antenna dark brown intermixed with few palebrown scales, flagellum dark brown; male with 4 th flagellum dorsally protuberant and curved, forming a deep concavity opposite slightly excavated flagellomeres 5–6 ; proboscis dark brown. Thorax: Tegula and mesonotum dark brown. Legs dark brown intermixed with few pale­brown scales to near apical margins of all segments and tarsomeres. Forewing (Fig. 337 ): Length 5.2 –7.0 mm (n = 15 ), dark brown intermixed with few brown and pale­brown scales; cell with one large, dark­brown spot near distal end. Undersurface dark brown. Hindwing: Pale brown basally, gradually darkening to brown towards apex. Abdomen (Fig. 150 ): Male with 4 transverse, irregular rows of sex scales on intersegmental membrane between terga 2–3 and terga 3–4 . Male Genitalia (Figs. 151–152 ): Uncus elongate, wide at base, broadly constricted medially, broadly rounded apically, slightly keeled dorsally, apex narrowly rounded; gnathos protuberant, narrow throughout length, upturned apically; tegumen slightly widened basally; dorsal strut of tegumen with arms convergent, fusing near midlength between posteroventral margin and base of uncus, dorsally divergent slightly and atrophied; costa of valva short, slightly sloped broadly from a shallow depression beyond a short, digitate process to dorsal furca; apex widely bifurcate, distance between furcae about as long as costa; dorsal furca curved outwardly, ventral furca twice as large as dorsal furca, and curved dorsolaterally; sacculus twisted apically slightly beyond midlength, constricting valva, widening distoventrally forming a broadly rounded cucullus; cucullus membranous and setose; vesica of aedeagus with many stout, conical cornuti, and bearing a large crescent­shaped, apical cornutus; cornutus with one side shortened near midlength, exposing a hollow, basal cavity. Female Genitalia (Fig. 281 ): Papillae anales elongate, slightly wider basally than apically; apophysis posterioris and apophysis anterioris about equal in size; eighth sternum slightly narrower than eighth tergum; eighth sternum with posterior and anterior margins broadly rounded, anterior margin more emarginate medially; eighth tergum with posterior and anterior margins broadly rounded; antrum enlarged posteriorly from a constricted anterior part; posterior margin nearly straight, posterolateral margin angular; anterior part with a pair of short, spinelike processes inside cavity formed from outer wall; ductus bursae short; corpus bursae denticulate, slightly enlarged anteriorly; ductus bursae large, coiled throughout length, coils gradually decreasing in size apically; ductus seminalis with a long, 8 ­coiled, internal support; side opposite ductus seminalis with an elongate, platelike, internal support. Type Examined: Holotype , ɗ, “Balboa, Panama , Canal Zone, at light, 19.6 .[ 19 ] 24 , St. George Expedn., C.L. Collenette.” “BM Slide No. 5645 .” [ BMNH ]. Other Specimens examined: ( 11 ɗ, 4 Ψ) 1 ɗ, “Est[ación] Las Pailas, 800 m , P[arque] N[acional] Rincón de la Vieja, Prov[incia] Guan[acaste], COSTA RICA , 24 Nov. a 26 Ene. 1993 , J. Sihezar, G. Rodriguez, L­N­ 306300 , 388600 ”, “ CRI 001, 211921 ”, “ɗ Slide No. 495 ”; 1 ɗ, “ 23 Oct. a 12 Nov. 1992 , C. Cano”, “ CRI 000, 920403 ”, “Slide No. 500 ”; 1 ɗ, 1 Ψ, “Agua Buena, P. N. Guanacaste, Prov. Guana., COSTA RICA , 200 m , 7–12 Feb. 1994 , E. Lopez, L­N­ 334800 , 364100, 2692”, “ CRI 001, 747346 ”, “Ψ Slide No. 496 ”; “ CRI 001, 747377 ”, “Slide No. 494 ”; 1 ɗ, “Quepos, 80 m , P. N. Manuel Antonio, Prov. Punt., COSTA RICA , R. Zuniga, Abr. 1991 , L­S­ 370900 , 448800 ”, “ CRI 000, 593398 ”, “Slide No. 618 ”; 1 ɗ, 2 Ψ, “Fca. Cafrosa, 1300 m , Est. las Mellizas, P. N. Amistad, Prov. Punt., COSTA RICA , M. Ramirez, Mar 1991 , L­S­ 316100 , 596100 ”, “ CRI 000, 355262 ”, “Slide No. 682 ”; “ CRI 000, 355393 ”, “Ψ Slide No. 684 ”; “ CRI 000, 355278 ”, “Ψ Slide No. 685 ”; 5 ɗ, 1 Ψ, Same label data as above except, “Abr.”, “ CRI 000, 475034 ”, “Slide No. 683 ”; 1 ɗ, “ CRI 000, 474676 ”; 1 ɗ, “ CRI 000, 474630 ”; 1 ɗ, “ CRI 000, 475017 ”; 1 ɗ, “ CRI 000, 475106 ”; 1 ɗ, “ CRI 000, 474553 ”; 1 ɗ, Same label data as above except, “ June 1991 ”, “ CRI 000, 663689 ”, ( 7 in INBio , 7 in USNM ). Distribution ( Map 27 ): In Costa Rica , Glyphidocera illiterata is known from four collecting sites from the northwestern part of the country, along the Cordillera de Guanacaste, southeast to the coast in south­central Costa Rica , east to the border near Panamá , along the Cordillera de Talamanca. G. illiterata is also known in Panamá from specimens described by Meyrick ( 1929 ) .