Revision of the taxonomy of Polycirrus Grube, 1850 (Annelida: Terebellida: Polycirridae)
Author
Glasby, Christopher J.
chris.glasby@nt.gov.au
Author
Hutchings, Pat
chris.glasby@nt.gov.au
text
Zootaxa
2014
2014-10-21
3877
1
1
117
journal article
5268
10.11646/zootaxa.3877.1.1
344cf70d-6a17-464b-9a35-40324bcef9d4
1175-5326
4948375
2695A2A6-2805-4FC6-B6B6-A8C68354B944
Polycirrus pumilis
Hartmann-Schröder, 1990
Fig. 53a–d
Polycirrus pumilis
Hartmann-Schröder, 1990: 77
, figs 40–43.
Type
locality.
South of Yamba
, NSW,
Australia
.
Material examined.
PARATYPES
: 2,
AM
W203315
,
Australia
,
New South Wales
,
Angourie Point
, south of
Yamba
,
29°21' S
154°22' E
,
17.i.1976
, intertidal
.
Description.
According to original description,
holotype
incomplete,
5 mm
long,
0.68 mm
wide for 27 segments.
Dorsum anteriorly smooth. Venter anteriorly with mid-ventral groove and discrete ventro-lateral pads; pads more-or-less smooth, extending from segment 2 to 8. Mid-ventral groove from segment 3.
FIGURE 52.
Polycirrus porcatus
Knox & Cameron.
Holotype NMV G1751. a. Anterior ventral view. b. Left side posterior lateral view of notopodium, chaetiger 4. c. Notochaeta from segment 7 illustrated at x40 and x100. d. Lateral and frontal views of uncini from segment 7. e. Lateral views of uncinus from mid-posterior body segment (figs d, e from
Hutchings & Glasby (1986)
; http://www.australianmuseum.net.au/Uploads/Journals/17666/185_complete.pdf
FIGURE 53.
Polycirrus pumilis
Hartmann-Schröder.
a. Head end, dorsal view. b. Head end, ventral view. c. Notochaetae. d. Anterior uncini. Redrawn from
Hartmann-Schröder 1990
; no scales on original.
Buccal tentacles numerous, of
one type
, cylindrical, uniformly thin and weakly grooved, some shorter than others, arising at junction between prostomium and upper lip. Prostomial ridge slightly curved, not extending laterally. Upper lip comprising single (medial) lobe only, margin of lobe straight; oral surface glandular, ciliated, with grooves leading to mouth. Inner lower lip oblong, narrow, smooth; outer region flat, shield-like, oblong, wider than long, smooth, extending posteriorly to segment 3. Achaetous segments visible dorsally but obscured by expanded outer lower lip ventrally (
Fig. 53a, b
).
Notochaetigerous segments 10–11, extending to segments 12–13. Notopodia digitiform, short, lobes similar, low and rounded. Notochaetae within a chaetiger consisting of
one type
, gradually elongating from dorsal to ventral, pinnate, posteriorly same form as those anteriorly (
Fig. 53c
). Neurochaetae beginning on segment 7, 20–24 uncini per row, more in posterior body segments. Uncini with medium length neck and concave base (
Type
2, as determined by morphometric study, see below), teeth above main fang arranged in double transverse series (MF:1–3:6–8) subrostral process present as low protuberance (
Fig. 53d
).
Nephridial papillae not visible in
paratype
.
Comments.
According to
Hartmann-Schröder (1990)
, nephridial papillae are present on chaetigers 1–3 of the
holotype
, that is, segments 3–5. Those on segments 3 and 4 would represent pre-gular membrane nephridial papillae as defined here. The ones on segment 5 would be post-gular membrane nephridial papillae.
The shape of the anteriormost uncini in
P. pumilis
is not typical of
Type
2 as defined by
Glasby & Glasby (2006)
, which has an elongate rostrum, long neck and arched concave base; however, the results of their morphometric analysis utilising all components of shape variation placed these uncini in the
Type
2 category. In addition, analysis of co-varying notochaetal characters by these authors provided further support for the grouping of this species with other species bearing
Type
2 uncini.
Polycirrus pumilis
probably belongs to a group of
Type
2 species having dimorphic uncini; these species also have early commencement of neuropodia, i.e., prior to segment 10.