Revision of Ommatius Wiedemann (Diptera: Asilidae). IV. Pygommatius subgen. nov. with twenty-five Afrotropical species Author Scarbrough, Aubrey G. Department of Biological Sciences, Towson University, Baltimore, MD 21252, USA; Author Marascia, Claudio G. text Zootaxa 2003 2003-07-04 228 1 1 94 http://biotaxa.org/Zootaxa/article/view/zootaxa.228.1.1 journal article 5623 10.11646/zootaxa.228.1.1 081ba61b-c79c-401c-a265-060a46979cd2 1175­5334 5019043 AC9F77DE-F702-4620-B756-D753E77FBC73 Ommatius (Pygommatius) porticus , sp. n. Figs. 18 , 45 , 189­196 Male. Brown. Length, body 10.7­12.0 mm; wing 7.0­ 7.8 mm . Head : Yellow to yellowishgray tomentose. Face sparse setose, setae mostly short, longer ventrally; 6 thin, brown and 4 pencil­like yellow bristles present; FHWR 1.0:10.0­1.0:10.5. Proboscis with narrow base yellow. Antenna brownish­yellow, flagellum darkest. Frons brown­yellow tomentose, brown setose. Ocellar tubercle with longest seta as long as pedicel and scape combined. Occiput with 5­6 brown postocular bristles, longest bristle about half distance toward ocellar tubercle. Thorax : Mesonotum largely dark brown, corners yellow laterally, yellowish­brown dorsally; tomentum mostly brown, yellowish­brown to yellow laterally and posteriorly; narrow paramedial stripe boarded by thin, yellowish­brown tomentum; setae sparse, mostly brown, 4 lateral and 2 dorsocentral brown bristles present. Scutellum dark brown, yellowish­gray tomentose, sparse short pale setae and 2 brown marginal bristles present. Mediotergite black. Pleuron mostly yellow, anepisternum, katepisternite and sometimes meron black; yellow to yellowish­gray tomentum and pale setae present. Halter yellow. Wing ( Fig. 18 ): Apical third and posterior two­thirds dense microtrichose. Cell r 4 triangular, vein R 5 slightly undulant. Apex of cell M 3 slightly curved, just beyond crossvein rm. Leg : Coxae and anterior 2 trochanters yellow; hind trochanter brown posteriorly, 1 short brown bristle present. Femora mostly yellow; fore and middle femora yellowishbrown to brown apically; apical third­half of hind femur brown, brown sometimes extends dorsally and ventrally to basal third. Fore femur (see O. brevicornis , Fig. 23 ) ventrally with row of 6­8 yellow bristles. Middle femur with 3 long, thin, yellow bristles anteriorly; only setae present ventrally, bristles absent. Hind femur with only 4­5 anterior brown bristles; anteroventral row with brown setae, about as long as vertical height of hind femur; 1 ventrobasal bristle present; abundant setae and 3­5 bristles present posteriorly; HFWLR 1.0:3.9­1.0:4.3. Tibiae yellow with narrow apices usually brown; fore and middle tibiae with long, thin and yellow bristles, longest about three­fourths as long as fore tibia. Hind tibia ( Fig. 45 ) mostly yellow, narrow apex and base brown and brownish­yellow respectively; shaft flattened medially before preapical constriction. Tarsi with basal tarsomere of fore and middle tarsi yellow; remaining tarsomeres light brown to brown; fore tarsus laterally with 1­2 yellow bristles. Abdomen: Mostly dark brown, wide lateral margins of tergites 1­4 and sternites 1­4 entirely yellow, with brown tomentum and brown setae dorsally; tergite 6 constricted, midapical margin of tergite 6 slightly produced, apical corner of tergite 3 with 2­3 and entire narrow lateral margin of tergite 4 with a fringe of long, yellow, sometimes brown and yellow mixed, bristles. Sternites 1­6 yellow to yellowish­brown; sternites 4­6 with abundant, erect, thin, yellow bristles; none exceptionally long or thick; sternite 7 mixed yellow and brown setose; sternite 8 brown setose. Terminalia ( Figs. 189­193 ): Ventral lamella with long median ridge, latter with abundant, long, thin, yellow setae basally and thicker brown setae apically. Epandrium 3 branched; middle and ventral branches capitate, the former slightly longer, the latter with 1 flat, leaf­like, and 4­5 hooked bristles. Gonostylus podiform. Gonocoxite laterally with minute setae arising from large sockets. Hypandrium without apical tuft of bristles. Female. Differs from male as follows. Length, body 9.0­ 10.5 mm ; wing 7.0­ 7.8 mm . Head : Bristles thinner than in male. FHWR 1.0:10.1­1.0:10.7. Mystax with yellow or black bristles. Leg: Normal, without unusual shapes of hind femur and hind tibia and unusual vestiture of fore and hind femora.; HFWLR 1.0:5.4­1.0:5.8. Abdomen : Without unusual vestiture and constriction in males. Tergites 1­7 usually with narrow lateral margin yellow; tergites 5­8 with an apical row of bristly setae, at least 1 in apical corner of tergite 4. Sternites with mostly yellow vestiture; sternites 5­7 apically with several bristly, brown setae, and laterally with 2­3 bristles. Tergite 9 membranous dorsally, extremely short. Terminalia ( Figs. 194­196 ): Three spermathecae present; duct basal. Sternite 8 with apex acutely triangular, with medioapical ridge that apically forms a small hooked spine; surface thin, membranous; several long, thick, brown bristles present. SPECIMENS EXAMINED. CÔTE D'IVOIRE : Holotype ♂ , allotype ♀ , Banco Nat. Park N. Abidjan / S. side 23­27.iv.89 / 05 o 22' N : 04 o 03' W / JGH Londt edge of wide tract / in forest ( NMSA ) . Paratypes : CÔTE D'IVOIRE : 1 ♂ , 9 ♀ , same data as holotype ( NMSA ) ; 1 ♀ , Asagny Nat. Park / roadway between gate and camp / 26.iv.89 05º14’N : 04º49’W / JGH Londt forest and forest edge ( NMSA ) ; 1 ♀ , Haute / Cerde de Mankono / entré Kourokoro and Mankono / ( F. Fleury ) / A. Chevalier / vi.10 ( MNHN ) ; 4 ♀ , Lamto / Gal. forestière / 24.x.70 / D. Lachaise ( MNHN ) ; 1 ♀ , Lamto / 13.x.70 / D. Lachaise ~ forêt galerie ( MNHM ) ; 1 ♀ , same data except 4.x.70 ~ Lisière / forêt galerie ( MNHN ) ; 1 ♀ ( Fernando­Poo ) Forêt / RTE de Concepcion ~ 4km . / Aprés Carref. De Moka Alt. 400 m / 13.xi . 70 L. Matile ( MNHN ) ; 3 ♂ , 1 ♀ , Fauch Quant / 16.i.71 / GFB / D. Lachaise / ( MNHN ) ; 2 ♀ , Fauch Quant / GFB / 25.ix.70 ( MNHN ) ; 1 ♂ , same data except 3.iv.71 ~ Lamto / D. Lachaise ( MNHN ) ; 1 ♀ , Fauch Quant / D1 VG / 7.xii.70 ( MNHN ) ; 1 ♀ , Lamto Bas Fond / Savane / non brûleé / 22.i.71 / D. Lachaise ( MNHN ) ; 4 ♂ , 4 ♀ , Marigot / Salé ~ 7.vi.71 ~ Museum Paris / Lamto / D. Lachaise ( MNHN ) ; 1 ♂ , same data except 19.v.71 ( MNHN ) ; 1 ♀ , same data except Pandanus 2 / 6.v.71 14h ( MNHN ) ; 1 ♀ , same data except Raphia­I / 8.v.71 ( MNHN ) ; 1 ♂ , same data except Zonaphia / 7.v.71 ( MNHN ) ; 1 ♀ , same data except J. P. Cola ( MNHN ) ; 2 ♂ , Marigot / Salé / 13.i.71, 15.i.71 ~ Lamto / D. Lachaise ( MNHN ) ; 2 ♂ , 1 ♀ , Marigot / Salé ~ 26.i.71 ~, Lamto / D. Lachaise ( MNHN ) ; 1 ♂ , Marigot / Salé / 15.i.71 ~ Lamto / D. Lachaise ( MNHN ) ; 1 ♀ , Marigot / Salé 9h / 3.iii.71, Lamto / D. Lachaise ( MNHN ) ; 1 ♂ , same data except rn­ny / 15.ii.71 ( MNHN ) ; 1 ♂ , Ficus capensis / fruits à terre / Lamto 7.i.71 / D. Lachaise ( MNHN ) ; 1 ♀ , Savane non / brûleé / bas depente / 7.i.71 / Zone Afromomum / bosquet / pres GFB / D. Lachaise ( MNHN ) ; 1 ♂ , Lamto / 6.vii.71 / D. Lachaise ~ Galerie ( MNHN ) ; 1 ♂ , 15.viii.72 / rive du Bandama Blac / radier de la route de / Tort a à Niakara­ mandou­ / gou, 5º32’W­ 8º46’N / J. Hamon ( MNHN ) ; 1 ♀ , Lisière / F. Plateau / Foro­ Bouaké / 24.vii.74 / G. Couturier ~ O. R .S. T .O.M. ( MNHN ) ; 1 ♀ Dabadougou / 16.xi.74 / G. Couturier , bord Marigot ~ O. R .S. T .O.M. ( MNHN ) ; 1 ♀ , Bouaké / Foro­Foro / 17.x.74 / G. Couturier ~ O. R .S. T .O.M. ( MNHN ) ; 1 ♂ , 1 ♀ , same data except foret galarie / M5B ( MNHN ) ; 1 ♀ , Gal. For. Band. / Fauch. Quant. / 9.xi.70 ( MNHN ) ; 3 ♀ , Adiopodumé / 27.iii.75 / G. Couturier ( MNHN ) ; 1 ♂ , 2 ♀ , same data except 1.xi.74, 7.iii.75, 4.iv.75 / G. Couturier Chasse , ( MNHN ) ; 1 ♀ , same data except Chasse / 23.i.75 ( MNHN ) ; 1 ♀ , Cascade / Mam / 21.xi.74/ G. Couturier ( MNHN ) ; 2 ♀ , Taï / 15­20.viii.78 / G. Couturier ~ forêt dense / sempervirente ( MNHN ) . GUINEA : 1 ♂ , Camp 4 ( 1000 m ), Museum Paris / Nimba ( Guinee ) / M. Lamette 11.vi.48 ( MNHN ) . NIGERIA : 1 ♀ , Lagos , G. Strachan ( BMNH ) ; 1 ♂ , 1 ♀ , T . Cacao ~ C. R .I.N. / 7:ll:1986 / Gambari / R . H. Booker ( BMNH ) ; 1 ♀ , Ikeja Airport / Lagos Nigeria / 23.xii.48 / B. Malkin ( CASC ) . Distribution. A forest edge species captured throughout the year in West Africa. Etymology. Latin, porticus , for ‘gallery or path’ [forêt galerie] in a forest. Remarks. In addition to the characters in the key, the male O. porticus is recognized by the combined characters of the terminalia ( Figs. 189­193 ) and the two rows of erect setae on sternites 4­6.