Phylogeny of Dynastor and Brassolis butterflies (Lepidoptera: Nymphalidae): a tough nut to crack Author Garzón-Orduña, Ivonne J. Author Penz, Carla M. text Zootaxa 2009 2134 1 22 journal article 10.5281/zenodo.188448 10d3b521-818c-44d2-af1b-df4281f82b80 1175-5326 188448 Genus Dynastor Doubleday Robust-bodied, ranging from medium ( darius FW length 97.9–109.8 mm, macrosiris 94.6 mm) to large ( napoleon 116.7–131.7 mm). Antennae mildly clubbed. Dorsal FW brown; postmedial band cream-white, always distal to the discal cell in both sexes, broken ( Fig. 1 ). Costal FW margin ripple-patterned in both sexes. Dorsal HW marginal line usually thin and cream-white (thick and bright orange in napoleon ); postmedial band usually absent (except darius ). Ventral surface of both wings either mottled or ripple-patterned ( macrosiris ). Ventral HW usually with three eyespots, in cells Sc+R1 (elongated), M1, and Cu1 (C-shaped). Extra eyespots may be present. Monomorphic for wing color and shape, males smaller than females. Male gnathos bifurcated, varying in shape between species; phallus short, with few small spines on the shaft; valva narrow apically, with a sclerotized costal margin bearing small to large spines ( Fig. 8 E). Female lamella antevaginalis absent ( Fig. 9 B), corpus bursa with paired signa.