New subgenus Monilistylus subgen. n. and a new species Procloeon (Monilistylus) ornatipennis sp. n. (Ephemeroptera: Baetidae: Procloeon) Author Kluge, Nikita J. text Zootaxa 2020 2020-02-21 4742 3 573 587 journal article 10.11646/zootaxa.4742.3.11 a1b47413-2f76-404c-86e4-571b06c6a1cd 1175-5326 3677917 B5B11009-059B-4DC1-85B6-831A67B72166 Cloeon /fg1 Branch «6»: Kluge & Novikova 1992 : Fig. 11 . Hierarchica name: Cloeon /fg1: Kluge 2012: 362 . Tribus Cloeonini: Kluge 2016: 492. Diagnosis of Cloeon /fg1. Larva. Both mandibles with dense setae between prostheca and mola ( Figs 4 , 18, 20 ). Labial palp with 3rd (distalmost) segment truncate ( Figs 26–27 ). On anterior side of tibia, proximal end of patella-tibial suture bent distally and connected under acute angle with row of long thin hairs ( Figs 30–31 ) (patella-tibial suture has equal structure on all legs – see Anteropatellata). Claw symmetric, slightly curved, with 2 similar rows of denticles ( Fig. 32 ); in some taxa without denticles. Abdominal segment IX and often one or several previous uromeres with longitudinal row of spines on each lateral margin ( Fig. 35 ). Tergalii often with anal-proximal expansion curved dorsally and forming a second (dorsal) lamella of tergalius; this lamella is absent in selected species of Procloeon /g1, including both species of Monilistylus ( Figs 9–15 ). Tergalii able to intensive rhythmic respiratory movements. Larval cerci and paracercus with dark rings on each 4th joining ( Fig. 8 ). Subimago. All tarsomeres covered with pointed microlepides, without blunt microlepides. Process of posterodorsal angle of postsubalar sclerite diminished, pointed, with concave antero-dorsal margin ( Fig. 43 ) (unlike other Turbanoculata, in which this process is larger and has convex antero-dorsal margin). Winged stages (imago and subimago). On fore wing each space with no more than 1 marginal intercalary ( Fig. 44 ). Hind wing, if present, of « Centroptilum - type »: relatively narrow and long, with hooked costal projection (in both species of Monilistylus hind wings absent). Male genitals . Gonovectes fused by their median ends with penial bridge, unable for protraction ( Fig. 52 ). Gonostylus with 2nd segment thickened apically, 3rd (distalmost) segment clavate ( Figs 49 , 53 ). In mature larva ready to molt to subimago, subimaginal gonostyli folded under larval cuticle by « Cloeon - type »: 2nd segment directed laterally, 3rd segment curved either caudally ( Fig. 34 ), or caudally-medially. FIGURES 1–8. Larval exuviae. 1–2, Procloeon ( Monilistylus ) monilistylus : 1, right half of pro- and mesonotum; 2, thoracic sterna, pleura, metanotum, abdominal sterna and terga. 3–8, Procloeon ( Monilistylus ) ornatipennis sp. n. : 3, right half of pro- and mesonotum; 4, head; 5, thoracic sterna, pleura, metanotum and two legs; 6, abdominal sterna and terga (with caudalii of holotype); 7–8, caudalii of holotype, enlarged. FIGURES 9–25. Details of larvae. 9–23, Procloeon ( Monilistylus ) ornatipennis sp. n. : 9–15, tergalii I–VII (holotype); 16, labrum; 17, maxilla (dorsal view); 18, right mandible; 19, its kinetodontium rotated after pressure of slide; 20, left mandible; 21–23, its incisor, kinetodontium and prostheca rotated after pressure of slide. 24–25, Procloeon ( Monilistylus ) monilistylus , kinetodontium and prostheca rotated after pressure of slide. Composition. Taxon Cloeon /fg1 includes Similicloeon Kluge & Novikova 1992 (= Intercloeon Kluge & Novikova 1992 ), Cloeon /fg2 (or genus Cloeon Leach 1815 s. str. ), Waynokiops Hill, Pfeiffer & Jacobus 2010 , Pseudocentroptilum Bogoescu 1947 [with the single species P. unguiculatum Tshernova 1941 (= P. motasi Bogoescu 1947 )] and Procloeon /g1 (see below).