New data on the Harpacteinae of Turkey (Araneae, Dysderidae) Author Kunt, Kadir Boğaç Department of Biology, Faculty of Science, Eskişehir Technical University, TR- 26470 Eskişehir, Turkey & Cyprus Wildlife Research Institute, Taşkent, Kyrenia, Cyprus Author Özkütük, Recep Sulhi Department of Biology, Faculty of Science, Eskişehir Technical University, TR- 26470 Eskişehir, Turkey text Zootaxa 2023 2023-11-23 5375 3 379 408 https://mapress.com/zt/article/download/zootaxa.5375.3.4/52335 journal article 10.11646/zootaxa.5375.3.4 1175-5326 10199980 26591537-2135-4301-AD46-FF4D2D9F7C68 Harpactea kankilicorum sp. nov. Figs 55–61 , 63–64 Material examined: Holotype 1 ♂ ( ETZM ), Kemer Dist. , Beetwen Sarısu-Beldibi ( 36°47’6.26”N 30°34’9.29”E ), asl c. 18 m , 25 Feb 2015 , leg. Kadir Boğac Kunt. Paratypes 1 ♀ ( ETZM ), same data as holotype — 3 ♂ , 3 ♀ ( ETZM ), Kemer Dist. , Göynük Canyon ( 36°41’0.02”N 30°31’36.95”E ), asl c. 104 m , 10 Mar 2016 , leg. Kadir Boğac Kunt — 2 ♀ , 1 ♂ , 4J ( ETZM ), Kumluca Dist. , Adrasan Vill. ( 36°17’47.22’’N 30°28’31.07’’E ), asl c. 20 m , 25 Feb 2016 , leg. Kadir Boğac Kunt — 4 ♀ , 1 ♂ , 4J ( ETZM ), Kemer Dist. , Gedelme Plateau ( 36°37’30.02”N 30°26’40.20”E ), asl c. 763 m , 25 Mar 2017 , leg. Kadir Boğac Kunt — 1 ♀ ( ETZM ), Kemer Dist. , Üçoluk Plateau ( 36°38’50.49”N 30°26’1.68”E ), asl c. 1078 m , 25 Mar 2017 , leg. Kadir Boğac Kunt. Etymology: The new species was named in honor of the “Kankılıç Family (Zeynel Abidin, Selvi, Tolga, Teoman and Gülbahar)”, which made very important contributions to the education in the life of the first author. FIGURES 55–56. Harpactea kankilicorum sp. nov. 55. Male, habitus 56. Chelicerae, ventral view (red dots, promarginal; white dots, retromarginal teeth). Diagnosis: The male palp of H. kankilicorum sp. nov. resembles H. alanyana Özkütük, Elverici, Marusik & Kunt, 2015 , H. cressa Brignoli, 1984 and H. sturanyi (Nosek, 1905) by the distal projection of the anterior part of the tegulum, the long, rounded embolus, bent 90°, and the sheet-like conductor with two distal apophyses. It is distinguished form the former species by the distal projection of the tegulum. Of these species, H. alanyana lacks the medial apophysis. The conductor consists of two prominent apophyses in H. alanyana , H. cressa and H. sturanyi , while in H. kankilicorum sp. nov. it is a double-layered plate ( Fig. 61 ). Also in H. kankilicorum sp. nov. , embolus curves distally more tightly than in H. alanyana ( Figs 57–62 ). The vulva of H. kankilicorum sp. nov. resembles that of H. nuragica Alicata, 1966 , an endemic of Sardinia island, by the short, and distaly rounded spermatheca, with a keel projected distally, but it can be differentiated by much more swollen spermatheca in H. kankilicorum sp. nov. Description: Measurements: [ Holotype ♂ / Paratypes ♀ ] TL 2.60/3.40–3.60; AL 1.45/2.15–2.25; CL 1.15/1.25–1.35; CW 1.10/1.10–1.20; Clh 0.03/0.02–0.02; AEd 0.07/0.08–0.08; iAE 0.02/0.02–0.02; PLEd 0.07/0.07–0.07; PMEd 0.05/0.06–0.06; ChF 0.17/0.20–0.23; ChG 0.17/0.20–0.23; ChL 0.36/0.51–0.55. [ Paratype ♂ ] TL 3.15; AL 1.65; CL 1.50; CW 0.95; Clh 0.02; AEd 0.09; iAE 0.02; PLEd 0.08; PMEd 0.07; ChF 0.18; ChG 0.18; ChL 0.47. Small sized harpacteine spiders. Carapace reddish in both sexes. Thoracic region pentagonal, head region roundish anteriorly. Transition between head and thoracic regions distinct. Fovea dark brown, longitudinal, distinct ( Fig. 55 ). AE, PLE and PME close to each other, arranged annularly. AEs, PLEs, and PMEs are seperated. Sternum, labium, gnatocoxae brown. Chelicerae reddish, brown. The promarginal part of the cheliceral groove is relatively strong; the retromarginal part has two very small teeth. The one at the base of the promarginal tooth is larger than the other, and there is an opening between the two, approximately the length of the base width of the first. The retromarginal tooth near the base of the cheliceral groove is aligned with the centre of the first of the promarginal teeth. The retromarginal teeth are six to seven times as far apart as their basal length ( Fig. 56 ). Abdomen cylindrical, white; covered with blackish setae. Legs brownish. Leg formula: VI, I, II, III. Leg spination and measurements given in tables 13 and 14.

Legs	I	II	III	IV
♂
C	0	0	0	2 pl
Fe	2 pl	(1) 2 pl	3 rl 1 d	1-1 d
Pa	0	0	1 rl	0
Ti	0	0	2 rl (1) 2 pl 1-1-2 v	1-1-1 rl 1-1-1 pl 1-1-2 v
Me	0	0	3 rl 2 pl 1-1-2 v	1-1-1-1 rl 1-1-1 pl 1-1-2 v
♀
C	0	0	0	(2) 3 pl
Fe	2 pl	(0) 1 d 1-1 pl	2 rl 1 d	(2) 3 d (0) 2 pl
Pa	0	0	1 rl	0
Ti	0	0	1-1 rl (1) 2 pl (3) 4 v	1-1-1 rl 1-1-1 pl 1-1-2 v
Me	0	0	1-1-1 rl (2) 3 pl (2) 4 v	1-1-1-1 rl 1-1-1 pl 1-1-2 v

Legs	I	II	III	IV
♂
Fe	0.94–0.99	0.88–0.94	0.64–0.70	1.05–1.28
Pa	0.59–0.64	0.47–0.47	0.35–0.41	0.53–0.63
Ti	0.85–0.91	0.76–0.82	0.41–0.59	0.94–1.00
Me	0.64–0.82	0.59–0.64	0.59–0.70	0.99–1.25
Ta	0.29–0.29	0.23–0.23	0.18–0.18	0.29–0.30
Total	3.31–3.65	2.93–3.10	2.17–2.58	3.80–4.46
♀
Fe	1.11–1.13	0.82–1.03	0.70–0.75	1.30–1.35
Pa	0.64–0.75	0.60–0.75	0.35–0.43	0.65–0.65
Ti	0.88–0.90	0.80–0.90	0.63–0.70	1.15–1.20
Me	0.75–0.76	0.68–0.75	0.80–0.80	1.25–1.28
Ta	0.28–0.30	0.28–0.30	0.28–0.30	0.26–0.38
Total	3.66–3.84	3.18–3.73	2.76–2.98	4.61–4.86

Palp ( Figs 57–61 ): Palpal tarsus is shorter than the tibia. Tegulum piriform ( Fig. 57 ). Palpal tarsus conical, shorter than tibia ( Fig. 60 ). Transition zone between tegulum and distal extensions is long and rectangular, strikingly ( Fig. 59 ). Embolus black and strongly sclerotized ( Fig. 57 ). Located on the posterior edge of this transition zone and there is a deep indentation just before it. The embolus originates at the distal end of this indentation, curving strongly towards the median apophysis as a hook. Conductor is a double-layered structure ( Fig. 61 ), bearing a slightly curved tip. Median apophysis patelliform, located just opposite the embolus base ( Fig. 61 ). FIGURES 57–62. Harpactea kankilicorum sp. nov. 57. Bulb (left), prolateral view 61. Ditto, SEM photo 58–59. Ditto, nearly retrolateral view 60. Ditto, retrolateral view (arrow indicates the deep indentation) 62. Harpactea alanyana , bulb (left), retrolateral view. Abbrevations: C , conductor: E , embolus; MA , Median apophysis; TZ , transition zone. Vulva ( Figs 63–64 ): The first quarter of the spermatheca is developed as spermathecal keel and is spinous. The second and third quarters expanded and took the shape of a sphere ( Fig. 63 ). The fourth quarter is rod-shaped and almost half the width of the spherical part. The ends of the anterior margin of the anterior arc are directed anteriorly. Transversal bar short. Posterior diverticulum membranous, circular ( Fig. 63 ).