The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina) Author Erwin, Terry L. text Zootaxa 2002 119 1 68 journal article 10.5281/zenodo.162803 f514e964-2196-461c-84ff-64243643bc81 1175­5326 162803 Agra catie Erwin , new species ( Figs. 7a, 7 b, 7c, 7d) Holotype . Male, COSTA RICA : Limón, Tortuguero National Park, Estacíon Cuatro Esquinas, sea level, 10°34'N 83°32'W , September (R. Delgado)( INBio : CRI 000­840274). Diagnosis . Femur concolorous. Prothorax narrow, not wider than head across eyes. Elytron with sutural apex markedly prolonged, acutely pointed. Male middle and posterior tibiae excavate and densely setose in apical half; elytral apex shallowly arcuate between lateral and apical apices; metasternum pubescent; abdominal sterna 2­5 with 2 bilateral patches. Description . Color and luster: Prothorax and elytra dark piceous; head and appendages dark rufous. Surface very shiny. Form: Head ( Fig. 7a ) behind eye moderately elongate, markedly tapered in male, less so in female. Male middle and posterior tibiae excavate and densely setose in apical half. Elytron ( Fig. 7 b) with apex shallowly arcuate between lateral and apical apices. Metasternum pubescent. Abdominal sterna 2­5 with 2 bilateral patches. Abdominal sternum VI ( Fig. 7 c) deeply V­notched in male, emarginate in female. Aedeagus ( Fig. 7 d) with ostium of medium length, apex simple cordiform, left lateral face of phallus with distinct impression; parameres glabrous. Size: medium, 15 to 19.6mm in length, 3.6 to 4.4mm in width. Other specimens examined. Costa Rica : 18 Paratypes : 5 males , 13 females from the following Conservation Areas: Cordillera Volcánica Central, Guanacaste, Osa, Pacifico Central, Tortuguero. Costa Rica , Cartago, 1f, Turrialba , 3.0 km SE , CATIE , 83° 38' W , 09° 53' N , June (E. Giesbert)( HAH : ADP4424), 1f, IICA / CATIE , April (E. Giesbert)( FSCA : ADP4421); 1m , Guanacaste, P.N. Guanacaste, Estacíon Pitilla , Santa Cecilia, 9.0 km S, 700m , 85° 25' 40' W, 10° 59' 26 N, LN 330200,380200, March (GNP Biodiversity Survey)( INBIO : CRI 000­034152), 1f, 85° 25' 40' W, 10° 59' 26 N, December (C. Moraga & P. Rios)( INBIO : CRI 000­166086); 1f, Heredia, Puerto Viejo, 3.0 km S, Finca La Selva, 84° 01' W , 10° 26' N , April (H. A. Hespenheide )( HAH : ADP5096), 2f, March (H. A. Hespenheide )( HAH : ADP62307, ADP80772); 1m , Limón, Finca de E. Rojas, Sector Cerro Cocori, 150m , May (E. Rojas)( INBIO : CRI 000­468171), 1f, September (E. Rojas)( INBIO : CRI 000­951292), 1f, Rio Reventazon, Ebene, Hamburg Farm, 10, 83° 28' W , 10° 15' N , March (F. Nevermann)(WAR: ADP3873, 1m, A.C. Tortuguero, Sardinas, Barra del Colorado, 15m , 83° 33' W , 10° 40' N , LN 291900,565900 #4639, April ­ May (F.V. Araya)( INBIO : CRI 002­169837), 1f, 50m , LN 291900,565900 #2916, May (F.V. Araya)( INBIO : CRI 001­917493), 1m , 15m , January ­ February (F.V. Araya)( INBIO : CRI 002­163901), 1m , P.N. Tortuguero, Tortuguero, Cerro Tortuguero, 0­100m , 83° 32' W , 10° 34' N , LN 285000,588000, November (J. Solano ( INBIO : CRI 000­ 141193 ), 1f, A.C. Tortuguero, R.N.F.S. Barra del Colorado, Rio Sardinas, 10m , July (F.V. Araya)( INBIO : CRI 000­690645), 1f, 0m, July (R.U. Chavarría)( INBIO : CRI 000­244568); 1f, Puntarenas, Peninsula de Osa, Rancho Quemado, 200m , LS 292500, 511000, November (M. Segura)( INBIO : CRI 000­962800), 1f, Res. Biol. Carara, Estacíon Quebrada Bonita, 50m , LN 194500,469850, August (R. Zuñiga )( INBIO : CRI 000­ 181297 ). Specific epithet. The specific epithet, catie , is used as a noun in apposition. It is the acronym of the forestry school at Turrialba , Centro Agronómico Tropical de Investigación y Enseñanza. Notes . This species ranges from sea level to 1100m altitude.