Taxonomic and nomenclatorial revision of the Neotropical genus Phaeoxantha Chaudoir (Coleoptera: Cicindelidae) Author Moravec, Jiří 0000-0001-5294-6410 Mendel University in Brno, Faculty of Forestry and Wood Technology, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. jirmor @ quick. cz; https: // orcid. org / 0000 - 0001 - 5294 - 6410 jirmor@quick.cz Author Dheurle, Charles 0000-0001-5294-6410 Mendel University in Brno, Faculty of Forestry and Wood Technology, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. jirmor @ quick. cz; https: // orcid. org / 0000 - 0001 - 5294 - 6410 & 5 place Jenson, F- 52200 Langres, France. charles. dheurle @ wanadoo. fr; https: // orcid. org / 0000 - 0002 - 2580 - 6962 Corresponding author & Mendel University in Brno, Faculty of Forestry and Wood Technology, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. jirmor @ quick. cz; https: // orcid. org / 0000 - 0001 - 5294 - 6410 jirmor@quick.cz text Zootaxa 2023 2023-12-14 5386 1 1 83 https://www.mapress.com/zt/article/download/zootaxa.5386.1.1/52481 journal article 10.11646/zootaxa.5386.1.1 1175-5334 10376586 9A5C0CC4-3D86-45BD-97FC-694A4E31A8B5 Phaeoxantha ( Phaeoxantha ) cruciata cruciata ( Brullé, 1837 ) ( Figs 142–157 ) Megacephala cruciata Brullé, 1837: 2 . Megacephala ( Phaeoxantha ) cruciata cruciata : Horn 1910: 137 Phaeoxantha cruciata : Horn 1923: 211 . Megacephala ( Phaeoxantha ) cruciata cruciata : Wiesner 1992: 45 . Phaeoxantha cruciata : Cassola & Pearson 2001: 11 . Phaeoxantha cruciata cruciata : Erwin & Pearson: 2008: 55. Phaeoxantha cruciata cruciata : Wiesner 2020: 30 . Type locality . Argentina : Province of Corientes . Type material . Lectotype (designated here) ♂ in MNHN , labelled: “Muséum Paris / Province de / Corientes / d’ Orbigny 1834” [ochre-tarnished, printed] // “illegible number and “34” [circular, handwritten, partly illegible] // “cruciata ” [green, handwritten] // “ Type ” [red, printed]. Other material examined . 3 ♂♂ , 1 ♀ in CDCL : “ Paraguay / Dpto Concepcion , Cororo / 20-XI-98” . 1 ♂ , 1 ♀ in CDCL : “ Entre Rios , 3-94 / Argentine , Pena Leg / Dheurle coll” . 1 ♂ in CDCL : “ Paraguay , 17-02-92/ Dpto San Pedro, Nueva / Germania Paso Carmona / Dheurle coll” . 1 ♂ in CCJM : “ Entre Rios / Colon , XII.1981 ” . 1 ♂ in MHCP , 1 ♀ in CCJM : “ Argentina / Colon , “ Entre Rios / 24.II.1976 ” . 1 ♀ in CCJM : “ Argentina / Entre Rios ” . 1 ♀ in CCJM : “ Argentinien / Entre Rios , Colon / X.1976 , Kirschenhofer ” . 1 ♂ in MKPC : “ Paraguay , dept. Concepcion / Estancia Estrella / S22°06´ , W 57°43´ / 27.-28.II.2014 ”, U. Drechsel ”. 1 ♀ in MKPC : Paraguay depto San Pedro / Laguna Blanca / S23° 50´ , W56°20´ , 15.-17.II.2014 , U. Drechsel ” . Differential diagnosis . Phaeoxantha ( P .) c. cruciata markedly resembles P . ( P .) bifasciata (= P. aequinoctialis sensu auctorum ), being often confused with it both in collections and literature. Nevertheless, it may be distinguished by its wider, notably shorter and stouter-side elytra; in addition, the posteromedian black area is almost consistently of a mutually “chiropteran-like” shape with narrow “wings” ( Figs 142 , 148, 150 , 155 ). Therefore, also females of P . ( P .) bifasciata , particularly those whose elytra are less attenuated posteriad and sometimes possess a similar yet wider black pattern ( Figs 194–195 ), may be distinguished (apart from other characters). Examined adults of P . ( P .) c. cruciata are 15.50–17.7 (LT 17.5) mm long, 6.70–7.30 (LT 7.30) mm wide. Distribution . The nominotypical subspecies of Phaeoxantha ( P .) cruciata is known from the large area of the Rio de la Plata Basin. Apart from the Argentinean province Corientes ( type locality), the other examined specimens come from the Argentinean province of Entre Rios. Wiesner (2000) listed it also from Paraguay (provinces of Concepción , Itapúa and San Pedro ). The illustrated male specimen from Conception ( Wiesner 2000 , fig. 4) is in accordance with the examined and above-listed specimens from Concepcion . On the other hand, the photograph of a male presented by Wiesner & Bandinelli (2014: 4) as “ P. cruciata cruciata ” shows in fact a male of P . ( P .) bifasciata . The authors were probably influenced by the records of P . ( P .) cruciata from Bolivian localities in the departments of Chuquisaca and Santa Cruz including Rio Seco by Person et al. (1999a), which were evidently based on misidentified specimens. Examined specimens (COSJ) from some of the localities from Machareti near Villamontes in the Bolivian department of Chuquisaca ( Figs 214, 219 ), as well as those from Rio Seco near Cabezas in the Bolivian department of Santa Cruz ( Figs 202–213, 220 , proved to be in fact P . ( P .) bifasciata ( Brullé, 1837 ) ( = aequinoctialis sensu Fleutiaux (1894) and sensu auctorum – partim ) – see under the taxon below.