The tropical African genus Morgenia (Orthoptera, Tettigoniidae, Phaneropterinae) with emphasis on the spur at the mid tibia Author Massa, Bruno Department of Agricultural, Food and Forest Sciences, University of Palermo, Viale Scienze Bd. 5 A, 90128 Palermo, Italy bruno.massa@unipa.it Author Heller, Klaus-Gerhard Grillenstieg 18, 39120 Magdeburg, Germany Author Warchalowska-Śliwa, Elzbieta Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17, 31515 Krakow, Poland Author Moulin, Nicolas Route de l'Ecole, 82, Hameau de Saveaumare, 76680 Monterolier, France text Deutsche Entomologische Zeitschrift 2018 2018-08-02 65 2 161 175 http://dx.doi.org/10.3897/dez.65.26693 journal article http://dx.doi.org/10.3897/dez.65.26693 1860-1324-2-161 196D0BD954A1481181D4B714B6B2BEEE 22E30A28F3F95FB0848C3A58737B1CE9 1343164 Morgenia hamuligera Karsch, 1890 Figs 1 , 9 , 13 , 20 , 27 , 37 , 38 Morgenia hamuligera Karsch, 1890a. Entom. Nachricht. 16: 263. Morgenia hamuligera Type locality: Kribi (Cameroon) (MfN, Berlin). Material examined. Cameroon , Kribi ( holotype ♂ ) (MfN) ; Central African Republic , Sangha Special Reserve , Epiphyte 2008 Expedition , Camp 2, 22-23.X.2008 (light) ( 2♂ ) , P. Annoyer; Central African Republic , Dzanga-Ndoki National Park , Ndoki , Lakes Region (light) 31.I.-23.II.2012 ( 10♂ ) , Sangha 2012 team; Mboki , 24.I.2012 ( 3♂ ) , Sangha 2012 team; 35 km S Bangui , env. Salanga , 28-31.XII.2008 , J. Halada ( 1♂ ) ; Gabon , Lope National Park , SEGC-CIRMF 12.III.2013 (light), N. Moulin ( 1♂ ) ; Ivory Coast , Tai Nat. Park , Res. Station 16-20.III.2017 (light), B. Massa ( 5♂ ) ; Ivory Coast , Tai Nat. Park , Res. Station 22.III-4.IV.2017 (light) ( 7♂ , 1♀ ) , P. Moretto (BMPC). Distribution. M. hamuligera was the first species described in the genus Morgenia , from Kribi (Cameroon) and later reported from Barombi Station (Cameroon) ( Karsch 1890a , 1890b ). Griffini (1906 , 1908 ) recorded it from Fernando Poo (Equatorial Guinea) and Umangi (Democratic Republic of Congo), Ebner (1943) from Fernando Poo and Massa (2013) from Central African Republic. It is here reported from Gabon and Ivory Coast. Remarks. The short description by Karsch (1890a) points out the presence of a blackish spot on the stridulatory area of the left tegmen; the shape of this spot is similar to that of M. rubricornis (see below), but its inner margin is more or less rectangular, not triangular, as in Morgenia rubricornis (Figs 9 , 10 , 13 , 14 ). The mirror of the right tegmen is wide (Fig. 9 ). Antennae may be yellowish or reddish. The pattern of the costal area of the tegmina has regularly spaced crossveins (Fig. 9 ). The stridulatory file is 3.5 mm long, arched and composed of ca. 80 dense and evenly spaced teeth in the proximal part, followed by ca. 70 more or less widely spaced teeth in the distal part (Fig. 20 ). The inner ventral spur of the mid tibiae is long and does not exceed the first tarsal article; its length is about 18-20% of the tibia length. On the outer ventral margin of the mid tibiae, three close short spines are present (Fig. 27 ). Cerci are very stout, club-shaped, the ratio tegmina length/width is 4.0 (Figs 37-38 , Table 1 ). Figures 1-4. Habitus of the male of Morgenia hamuligera ( 1 ), M. rubricornis ( 2 ), M. plurimaculata sp. n. ( 3 ) and M. lehmannorum sp. n. ( 4 ) (left side for the first three species, right side for the fourth species). Table 1. Measurements of tegmina length and width and ratio of tegmina length/width of the eight species currently known in the genus Morgenia .

Species	Tegmina length	Tegmina width	Tegmina length/width
Morgenia hamuligera (n = 10)1	33.2 +/- 1.4 (30.3-34.9)	8.3 +/- 0.4 (8.0-9.2)	4.0 +/- 0.3 (3.5-4.2)
Morgenia rubricornis (n = 10)	29.4 +/- 1.1 (27.5-30.2)	7.5 +/- 0.5 (7.0-8.5)	3.9 +/- 0.2 (3.3-4.2)
Morgenia spathulifera (n = 5)	29.6 +/- 1.4 (27.9-31.4)	4.5 +/- 0.1 (4.3-4.6)	6.6 +/- 0.2 (6.4-6.8)
Morgenia plurimaculata sp. n. (n = 10)	34.1 +/- 0.9 (33.0-35.7)	8.0 +/- 0.2 (7.8-8.5)	4.2 +/- 0.1 (4.1-4.4)
Morgenia lehmannorum sp. n. (n = 1)	32.5	5.4	6.0
Morgenia angustipinnata sp. n. (n = 4)	32.0 +/- 1.7 (30.5-33.9)	5.0 +/- 0.4 (4.7-5.4)	6.4 +/- 0.1 (6.3-6.5)
Morgenia melica (n = 10)	34.3 +/- 2.2 (29.0-36.9)	7.6 +/- 0.3 (7.0-8.0)	4.5 +/- 0.2 (4.1-4.8)
Morgenia modulata (n = 10)	30.4 +/- 0.9 (29.1-31.4)	6.7 +/- 0.5 (6.0-7.6)	4.5 +/- 0.2 (4.1-4.9)

1 Massa (2017a) reports 5.5-5.6 as ratio length/width of tegmina in M. hamuligera and M. rubricornis , but more accurate measurements allowed to change them to 4.0 and 3.9, respectively.