A review of the tribes of Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae) Author Zahniser, James N. C44D6E44-FA1C-4B29-B7BB-FAF5940CD225 Illinois Natural History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 S. Oak St., Champaign, IL 61820, United States of America & urn: lsid: zoobank. org: author: C 44 D 6 E 44 - FA 1 C- 4 B 29 - B 7 BB-FAF 5940 CD 225 & corresponding author e-mail: zahniser @ illinois. edu Author Dietrich, Chris H. 82FCB86C-54B4-456A-AE5E-D7847D271CB9 Illinois Natural History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 S. Oak St., Champaign, IL 61820, United States of America & urn: lsid: zoobank. org: author: 82 FCB 86 C- 54 B 4 - 456 A-AE 5 E-D 7847 D 271 CB 9 text European Journal of Taxonomy 2013 2013-05-29 45 1 211 journal article 22221 http://dx.doi.org/10.5852/ejt.2013.45 a66592b8-63a8-4c2d-9471-e58ddb2c0559 2118-9773 3822710 41B10E4D-7DAB-40CA-A8FE-4ECA078E04A3 Drakensbergenini Linnavuori, 1979 Fig. 21 Type genus: Drakensbergena Linnavuori, 1961 . Diagnosis Drakensbergenini are small to large, ivory, ochraceous, to dark brown brachypterous leafhoppers, often with a dark brown median stripe from apex of crown to abdomen. They can be identified by the produced head, ocelli on the crown, tumid frontoclypeus, strongly tapering clypellus, and brachypterous forewings. Description HEAD. Head subequal to or wider than pronotum; somewhat to strongly produced. Discal portion of crown shagreen. Anterior margin of head shagreen or foliaceous; often shagreen and rounded to face laterally and foliaceous apically. Frontoclypeus tumid; texture shagreen. Clypellus strongly tapering apically; apex following or slightly surpassing normal curve of gena. Lorum distinctly narrower than clypellus near base. Antennal bases near upper or anterodorsal corners of eyes. Antennae short, less than 1.5 x width of head or long, 1.5 x width of head or longer. Gena strongly incised laterally (nearly forming right angle); with fine erect seta beside laterofrontal suture. Antennal ledge weakly developed (carinate or weakly carinate). Ocelli reduced or present; distant from eyes; on crown. Fig. 21. Drakensbergenini. A–B . Drakensbergena deorsuspina Stiller, 2009 . C . Drakensbergena sp. D–I . D . phaeogramma Stiller, 2009 . A–I . Standard views (see Material & Methods). THORAX. Pronotum lateral margin carinate or rarely ecarinate; lateral margin shorter than basal width of eye. WINGS. Forewing brachypterous; coriaceous; wings usually fused; veins obscure. LEGS. Profemur with AM 1 absent or reduced; intercalary row reduced or absent; row AV with relatively long macrosetae. Protibia dorsal surface rounded, convex. Metafemur apex macrosetae 2+1 or 2+1+1. Metatarsomere I expanded apically; plantar setae simple, tapered. MALE GENITALIA. Valve articulated with pygofer; lateral margin short, articulating with pygofer at a point. Pygofer basolateral membranous cleft present; macrosetae absent or reduced (≤ two rows). Subgenital plates free from each other; articulated with valve; without macrosetae. Style broadly bilobed basally, median anterior lobe pronounced. Basal processes of the aedeagus/connective absent or reduced. Aedeagus with single shaft and gonopore; apex bifid. Connective anterior arms somewhat divergent, Y- shaped; stem often appearing to be formed by two longitudinal bars; articulated with aedeagus. Segment X long; sclerotized dorsally and laterally. FEMALE GENITALIA. Pygofer with macrosetae reduced or absent. Ovipositor not protruding far beyond pygofer apex. First valvula not strongly convex; dorsal sculpturing pattern concatenate, reticulate, granulose, or maculose; sculpturing reaching dorsal margin or submarginal; without distinctly delimited ventroapical sculpturing. Second valvula abruptly broadened medially or subapically or broad, gradually tapered; without dorsal median tooth; dorsal teeth on apical 1/3 or more; teeth small, regularly or irregularly shaped. Third valvula with two irregular setal rows along entire ventral margin. Geography and ecology Distribution: Afrotropical region ( South Africa and Lesotho ). Drakensbergena spp. feed on grasses in the high elevation grassland biome of southern Africa. Remarks Drakensbergenini contains the type genus and 18 species.This group was considered a separate subfamily by Linnavuori (1979a) but placed as a tribe of Deltocephalinae by Zahniser & Dietrich (2010) . Stiller (2009) described 16 new species of the genus. Phylogenetic analyses and morphological characters (produced head, profemur intercalary row reduced, AM 1 seta reduced or absent, AV setae relatively long, metafemur apical setae 2+1+1 or 2+1) suggest that this tribe is related to Eupelicini. Selected references Linnavuori (1979a) , Stiller (2009) . Included genera Drakensbergena Linnavuori, 1961