Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species Author Vilkamaa, Pekka Author Menzel, Frank text Zootaxa 2019 2019-09-06 4665 1 1 67 journal article 21114 10.11646/zootaxa.4665.1.1 8053cc5c-900d-490b-84e9-1ca29a46aa3c 1175-5326 4000553 78F3F422-DE06-4C61-AC47-69AA684EB5AB Trichocoelina janetscheki ( Lengersdorf, 1953 ) comb. n. Figs 10 A, 10 B , 17 D Literature . Neosciara janetscheki Lengersdorf—Lengersdorf (1953) : 167, fig. 1; Janetschek (1956) : 471. Lycoriella ( Hemineurina ) janetscheki (Lengersdorf) — Tuomikoski (1959a) : 35; Gerbachevskaja-Pavluchenko (1986) : 31; Franz (1989) : 14; Menzel & Mohrig (2000) : 410; Wirta et al. (2016): appendix, unpaginated p. 21 (table S1) and unpaginated p. 39 (cladogram). Material studied . CANADA , Nunavut , Sverdrup Islands , Elles Ringnes Island , Isachsen , 70.79°N , 103.55°W , lemming burrow, 17.VI.1960 , J.R. Vockeroth , 4 males ( 3 in CNC, 1 in MZH ) ; same locality but moss near very small stream, 14.VII.1960 , J.F. McAlpine , 2 males (in CNC ) ; Nunavut , Ellesmere Island , Fosheim Peninsula , Hot Weather Creek , 79°58’N , 84°28’W , 2.VII.1990 , F. Brodo , 1 male (in MZH ) ; NE Greenland , Mestersvig , 72.24°N , 23,92°W , C. Vibe (in ZMUC ) ; NE GREENLAND , Zackenberg , 74°28’N , 20°34’W (UTM8265500:0513778), 44 m , 2–11.VII.2011 , T. Roslin & G. Várkonyi , 1 male (in MZH ) ; same locality but UTM 8265758:0513786, 37 m , Malaise trap , 11–20.VII.2011 , T. Roslin & G. Várkonyi , 2 males (in MZH , SDEI) ; same locality but 74.50°N , 21.00°W , 44 m , 7.VII.2011 , T. Roslin & G. Várkonyi , 1 male (BOLD Sample ID GRPV16, in DAUH) ; same locality but 74.4667°N , 20.5667°W , 48 m , 24.VII.2011 , T. Roslin & G. Várkonyi , 3 males (BOLD Sample IDs ZA2012- 50010, ZA2012-50011 and ZA2012-50012, in DAUH) ; USA , Colorado , Mt. Evans , 14 000 ft, 25.VII.1961 , W.R.M. Mason , 1 male (in USNM ) . Redescription . Male. Head . Face brown, antenna dark brown maxillary palpus pale yellowish. Eye bridge 2–3 facets wide. Face with 15–21 setae. Clypeus with 1–4 setae. Maxillary palpus with 3 segments, 1 st segment as long as or longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 3–9 setae, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.95–2.75x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere. Thorax . Dark brown, setae pale. Anterior pronotum with 5–11 setae. Proepisternum with 7–24 setae. Scutellum with 4 longer and some short and fine setae. Wing . Fumose. Length 2.2–3.1 mm . Width/length 0.35–0.40. Anal lobe weak. Veins distinct. c/w 0.50–0.60. R 1 /R 0.50–0.90. stM shorter than fork of M. r-m longer than bM, bM non-setose, r-m non-setose or with 1–2 setae. Halter yellow. Legs . Yellow, coxal setae pale. Fore tibial organ forming a large patch in shallow depression. Fore tibial spur as long as the tibial width. Abdomen . Pale brown, setae pale, short and fine. Hypopygium ( Fig. 10 B ). Brown, as abdomen. Intergonocoxal area long, with two short setose lobes. Gonocoxa broad, as long as gonostylus, medial margin basally smoothly curved, with short and fine setosity. Gonostylus ( Fig. 10 A ) voluminous, apically truncate, impressed; with short setosity, a short apical tooth, and numerous megasetae in the medial impression; megasetae slightly curved or straight, short and slender; with 1 well-differentiated whiplash seta basally at ventromedial margin. Tegmen ( Fig. 17 D ) longer than broad, laterally straight, apically with a narrow hyalinous acuminate process, weakly sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN . BOLD:ACK5495. Discussion . In Fig. 10 A the gonostylus is flattened and distorted to show the arrangement of the megasetae. In its extremely voluminous gonostylus, Trichocoelina janetscheki ( Lengersdorf, 1953 ) resembles most T. incrassata sp. n. , for distinguishing characters, see under the latter.