<document id="38B5740A782D7BB0AE647AAB6A14219A" ID-DOI="10.3897/zookeys.7.117" ID-GBIF-Dataset="7a12aaad-a568-4920-91cf-af3e9407eadc" ID-ISSN="1313–2970" ID-Zenodo-Dep="576436" ID-ZooBank="67B4D2EC-2C6D-4226-847D-0BA2B2777AE3" IM.materialsCitations_approvedBy="admin" IM.metadata_approvedBy="admin" checkinTime="1587986763134" checkinUser="plazi" docAuthor="Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; VandenSpiegel, Didier" docDate="2009" docId="1D3687F8A74DA64FFF62FCBFFD727166" docLanguage="en" docName="ZK_article_1970.pdf" docOrigin="ZooKeys 7 (7)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@60a754c6" docStyleName="zookeys.2008.journal_article" docTitle="Helodesmus Cook 1896" docType="treatment" docVersion="6" lastPageNumber="44" masterDocId="E10FFF80A766A664FFEEFF9FFF8E742C" masterDocTitle="Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="44" updateTime="1693518122439" updateUser="admin" zenodo-license-document="CC-BY-3.0">
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<mods:title id="7D2D0E5F8A0BE2326872DF4BD814CD7B">Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="502664BEBDD7E312C42D544571118FC4">Golovatch, Sergei</mods:namePart>
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<mods:affiliation id="B8ACA4364AFF2231DB2E7D0ACFA81A00">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<mods:namePart id="EA8B115792F6D95C12AA35D957B4F1CB">Geoffroy, Jean-Jacques</mods:namePart>
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<mods:affiliation id="DB68E23322962E75A5694FE55B3EAB77">Muséum national d’Histoire naturelle, Département Ecologie &amp; Gestion de la Biodiversité, UMR 7179 du CNRS, Equipe EVOLTRAIT, 4, avenue du Petit Château, F- 91800 Brunoy, France</mods:affiliation>
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<mods:namePart id="846EDE232DD69732E6C07FEE2AC38D15">Mauriès, Jean-Paul</mods:namePart>
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<mods:affiliation id="818F79C02631D9ECAFCA7015B1076901">Muséum national d’Histoire naturelle, Département Systématique et Evolution, USM 602, Section Arthropodes Case Postale n ° 53, 61 rue Buffon F- 75231 Paris, France</mods:affiliation>
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<mods:namePart id="891B695467B21B2329CB4DFA17CD5DE1">VandenSpiegel, Didier</mods:namePart>
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<mods:affiliation id="D6F5183C3F075674377A7BC998B5D01E">Musée Royal de l’Afrique centrale, B- 3080 Tervuren, Belgium</mods:affiliation>
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<mods:date id="DA79C20F8E775C9F39557916635ED09F">2009</mods:date>
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<treatment id="1D3687F8A74DA64FFF62FCBFFD727166" ID-DOI="http://doi.org/10.5281/zenodo.3792476" ID-GBIF-Taxon="163750768" ID-Zenodo-Dep="3792476" LSID="urn:lsid:plazi:treatment:1D3687F8A74DA64FFF62FCBFFD727166" httpUri="http://treatment.plazi.org/id/1D3687F8A74DA64FFF62FCBFFD727166" lastPageNumber="44" pageId="43" pageNumber="44">
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<paragraph id="952036EEA74DA64FFF62FCBFFD8F7716" blockId="43.[140,513,800,827]" box="[140,513,800,827]" pageId="43" pageNumber="44">
<heading id="CE688182A74DA64FFF62FCBFFD8F7716" box="[140,513,800,827]" fontSize="11" level="2" pageId="43" pageNumber="44" reason="3">
<emphasis id="A7EBEAFCA74DA64FFF62FCBFFD8F7716" bold="true" box="[140,513,800,827]" pageId="43" pageNumber="44">
Genus 
<taxonomicName id="529F4D6DA74DA64FFF31FCBFFD8F7716" ID-CoL="7PJQC" authority="Cook, 1896" authorityName="Cook" authorityYear="1896" box="[223,513,800,827]" class="Diplopoda" family="Haplodesmidae" genus="Helodesmus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="A7EBEAFCA74DA64FFF31FCBFFEE27717" bold="true" box="[223,364,800,827]" italics="true" pageId="43" pageNumber="44">Helodesmus</emphasis>
Cook, 1896
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<typeStatus id="4A24884CA74DA64FFF62FCF8FF4D77AD" box="[140,195,871,897]" pageId="43" pageNumber="44">Type</typeStatus>
species: 
<taxonomicName id="529F4D6DA74DA64FFEC8FCF8FD0577AD" authority="Cook, 1896" authorityName="Cook" authorityYear="1896" box="[294,651,871,897]" class="Diplopoda" family="Haplodesmidae" genus="Helodesmus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="porosus">
<emphasis id="A7EBEAFCA74DA64FFEC8FCF8FE7777AC" box="[294,505,871,897]" italics="true" pageId="43" pageNumber="44">Helodesmus porosus</emphasis>
Cook, 1896
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, by monotypy (
<typeStatus id="4A24884CA74DA64FFCD0FCF8FCFE77AD" box="[830,880,871,897]" pageId="43" pageNumber="44">type</typeStatus>
locality: Java, Indonesia).
</paragraph>
</subSubSection>
<subSubSection id="DD856565A74DA64FFF62FC31FCDC701E" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="952036EEA74DA64FFF62FC31FD7577C7" blockId="43.[140,1108,871,1074]" pageId="43" pageNumber="44">
= 
<taxonomicName id="529F4D6DA74DA64FFF4CFC31FE4A77EB" authority="Attems, 1930" authorityName="Attems" authorityYear="1930" box="[162,452,941,968]" class="Diplopoda" family="Haplodesmidae" genus="Gonomastis" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="A7EBEAFCA74DA64FFF4CFC31FE9177E4" box="[162,287,942,968]" italics="true" pageId="43" pageNumber="44">Gonomastis</emphasis>
<bibRefCitation id="F10E4B1FA74DA64FFEC9FC32FE4A77EB" author="Attems" box="[295,452,941,967]" firstAuthor="Attems" pageId="43" pageNumber="44" pagination="115 - 182" refId="ref18593" refString="Attems C (1930) Myriopoden von Java, Sumatra und Bali. Archiv fur Hydrobiologie, Supplement-Band 8: 115 - 182." type="journal article" year="1930">Attems, 1930</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="4A24884CA74DA64FFE21FC31FD8877E4" box="[463,518,942,968]" pageId="43" pageNumber="44">Type</typeStatus>
species: 
<taxonomicName id="529F4D6DA74DA64FFD87FC31FC6877EB" authority="Attems, 1930" authorityName="Attems" authorityYear="1930" box="[617,998,941,968]" class="Diplopoda" family="Haplodesmidae" genus="Gonomastis" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="A7EBEAFCA74DA64FFD87FC31FCCC77EB" box="[617,834,941,968]" italics="true" pageId="43" pageNumber="44">Gonomastis parvula</emphasis>
<bibRefCitation id="F10E4B1FA74DA64FFCA4FC32FC6877EB" author="Attems" box="[842,998,941,967]" firstAuthor="Attems" pageId="43" pageNumber="44" pagination="115 - 182" refId="ref18593" refString="Attems C (1930) Myriopoden von Java, Sumatra und Bali. Archiv fur Hydrobiologie, Supplement-Band 8: 115 - 182." type="journal article" year="1930">Attems, 1930</bibRefCitation>
</taxonomicName>
, by original designation. Synonymized by 
<bibRefCitation id="F10E4B1FA74DA64FFDD9FC4FFD7977C7" author="Hoffman" box="[567,759,976,1003]" firstAuthor="Hoffman" pageId="43" pageNumber="44" pagination="232 - 234" refId="ref19286" refString="Hoffman RL (1964) A note on the millipede genus Helodesmus (Polydesmida: Helodesmidae). Entomologische Berichten 24: 232 - 234." type="journal article" year="1964">Hoffman (1964)</bibRefCitation>
.
</paragraph>
<paragraph id="952036EEA74DA64FFF62FC6BFCDC701E" blockId="43.[140,1108,871,1074]" pageId="43" pageNumber="44">
= 
<taxonomicName id="529F4D6DA74DA64FFF4DFC6BFE6A7022" authority="Chamberlin, 1945" authorityName="Chamberlin" authorityYear="1945" box="[163,484,1011,1038]" class="Diplopoda" family="Haplodesmidae" genus="Porauxus" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="A7EBEAFCA74DA64FFF4DFC6BFE8B7021" box="[163,261,1012,1037]" italics="true" pageId="43" pageNumber="44">Porauxus</emphasis>
<bibRefCitation id="F10E4B1FA74DA64FFEE0FC6CFE6A7022" author="Chamberlin" box="[270,484,1011,1038]" firstAuthor="Chamberlin" pageId="43" pageNumber="44" pagination="1 - 43" refId="ref18759" refString="Chamberlin RV (1945) On some diplopods from the Indo-Australian Archipelago. American Museum Novitates 1282: 1 - 43." type="journal article" year="1945">Chamberlin, 1945</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="4A24884CA74DA64FFE01FC6BFDA87022" box="[495,550,1012,1038]" pageId="43" pageNumber="44">Type</typeStatus>
species: 
<taxonomicName id="529F4D6DA74DA64FFD65FC6BFBC07022" authority="Chamberlin, 1945" authorityName="Chamberlin" authorityYear="1945" box="[651,1102,1011,1038]" class="Diplopoda" family="Haplodesmidae" genus="Porauxus" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pangrangus">
<emphasis id="A7EBEAFCA74DA64FFD65FC6BFCFE7021" box="[651,880,1012,1037]" italics="true" pageId="43" pageNumber="44">Porauxus pangrangus</emphasis>
<bibRefCitation id="F10E4B1FA74DA64FFC97FC6CFBC07022" author="Chamberlin" box="[889,1102,1011,1038]" firstAuthor="Chamberlin" pageId="43" pageNumber="44" pagination="1 - 43" refId="ref18759" refString="Chamberlin RV (1945) On some diplopods from the Indo-Australian Archipelago. American Museum Novitates 1282: 1 - 43." type="journal article" year="1945">Chamberlin, 1945</bibRefCitation>
</taxonomicName>
, by original designation. Synonymized by 
<bibRefCitation id="F10E4B1FA74DA64FFD63FB88FCC0701E" author="Hoffman" box="[653,846,1047,1074]" firstAuthor="Hoffman" pageId="43" pageNumber="44" pagination="232 - 234" refId="ref19286" refString="Hoffman RL (1964) A note on the millipede genus Helodesmus (Polydesmida: Helodesmidae). Entomologische Berichten 24: 232 - 234." type="journal article" year="1964">Hoffman (1964)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DD856565A74DA64FFF62FBC2FD727166" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="952036EEA74DA64FFF62FBC2FD727166" blockId="43.[140,1108,1117,1355]" pageId="43" pageNumber="44">
<emphasis id="A7EBEAFCA74DA64FFF62FBC2FE85705B" bold="true" box="[140,267,1117,1143]" pageId="43" pageNumber="44">
<emphasis id="A7EBEAFCA74DA64FFF62FBC2FE88705B" box="[140,262,1117,1143]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Diagnosis</emphasis>
.
</emphasis>
Body with 19 segments, subcylindrical, not capable of volvation, with paraterga 2 rather well developed, but subsequent ones mostly represented by lateral swellings. Collum still large, covering the head from above, with 4-5 transverse rows of setigerous tubercles or pits. Following paraterga with three rows of similar tubercles or pits. Pore formula: 5, 7-17(18). Gonopods with poorly setose gonocoxae and a considerably shortened prefemoral+femoral (= setose) part, deeply biramous thereafter, with a long and flagelliform solenomere devoid of a hairpad.
</paragraph>
</subSubSection>
</treatment>
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