<document id="2F9A0D67F9EB762315D1E255FE4CFE17" ID-DOI="10.1649/0010-065X-75.3.537" ID-ISSN="1938-4394" ID-Zenodo-Dep="13251717" ID-ZooBank="urn:lsid:zoobank.org:pub:FCC13385-7D46-4918-B46F-5F344D5564C3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723001473221" checkinUser="felipe" docAuthor="Kamiński, Marcin Jan, Lumen, Ryan, Müller, Ruth, Kanda, Kojun, Robiche, Gérard &amp; Smith, Aaron Dennis" docDate="2021" docId="03D9ED45FFF74A2BFD06FA39F3D8621B" docLanguage="en" docName="TheColeopBull.75.3.537-578.pdf" docOrigin="The Coleopterists Bulletin 75 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-75.3.537" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Aptila costata Fahraeus 1870" docType="treatment" docVersion="2" lastPageNumber="551" masterDocId="FFE0953DFFF94A24FFABFFADF334635A" masterDocTitle="Illustrated Catalog of the Subtribe Helopinina Lacordaire (Coleoptera: Tenebrionidae: Blaptinae: Pedinini)" masterLastPageNumber="578" masterPageNumber="537" pageNumber="550" updateTime="1723004189327" updateUser="ExternalLinkService">
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<mods:title id="3E2F2E93AC2735B967567200F57947C4">Illustrated Catalog of the Subtribe Helopinina Lacordaire (Coleoptera: Tenebrionidae: Blaptinae: Pedinini)</mods:title>
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<mods:namePart id="3B07EF1CC0CD589780A467F5086D2C00">Kamiński, Marcin Jan</mods:namePart>
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<mods:namePart id="C5B9AC3BC5615A4D38A5E02366243F7E">Lumen, Ryan</mods:namePart>
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<mods:namePart id="8730AF8F75D297AEB07B4948022A2EF3">Müller, Ruth</mods:namePart>
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<mods:namePart id="7D400051F692A305A5B71489B84B2E0C">Kanda, Kojun</mods:namePart>
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<mods:namePart id="CD35377AE67EED4C9C08623A5A9DBA46">Robiche, Gérard</mods:namePart>
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<mods:date id="819379AE54A41AF80CB7B75B5899E20C">2021</mods:date>
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1. 
<taxonomicName id="4C7027D0FFF74A2AFD6EFA39F0DB66F3" authority="Fahraeus, 1870" authorityName="Fahraeus" authorityYear="1870" box="[709,1007,1427,1450]" class="Insecta" family="Tenebrionidae" genus="Aptila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="550" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="B9048041FFF74A2AFD6EFA39F07866F0" bold="true" box="[709,844,1428,1450]" italics="true" pageId="14" pageNumber="550">Aptila costata</emphasis>
<bibRefCitation id="EFE121A2FFF74A2AFCF8FA3EF0DB66F3" author="Fahraeus, O. I." box="[851,1007,1427,1449]" pageId="14" pageNumber="550" pagination="243 - 358" refId="ref25701" refString="Fahraeus, O. I. 1870. Coleoptera Caffrariae, annis 1838 - 1845 a J. A. Wahlberg collecta. Heteromera de- scripsit. Ofversigt af Kongliga Vetenskaps-Akad- emiens Forhandlingar 27: 243 - 358." type="journal article" year="1870">Fåhraeus, 1870</bibRefCitation>
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<treatmentCitationGroup id="AB607B7DFFF74A2AFD06FA02F0CA669F" box="[685,1022,1455,1477]" pageId="14" pageNumber="550">
<treatmentCitation id="0AD17A42FFF74A2AFD06FA02F0CE669F" author="Fahraeus, O. I." box="[685,1018,1455,1477]" page="259" pageId="14" pageNumber="550" year="1870">
<taxonomicName id="4C7027D0FFF74A2AFD06FA02F0CE669F" authority="Fahraeus 1870: 259" authorityName="Fahraeus" authorityPageNumber="259" authorityYear="1870" box="[685,1018,1455,1477]" class="Insecta" family="Tenebrionidae" genus="Aptila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="550" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="B9048041FFF74A2AFD06FA02F007669F" box="[685,819,1455,1477]" italics="true" pageId="14" pageNumber="550">Aptila costata</emphasis>
<bibRefCitation id="EFE121A2FFF74A2AFC92FA02F0CE669F" author="Fahraeus, O. I." box="[825,1018,1455,1477]" pageId="14" pageNumber="550" pagination="243 - 358" refId="ref25701" refString="Fahraeus, O. I. 1870. Coleoptera Caffrariae, annis 1838 - 1845 a J. A. Wahlberg collecta. Heteromera de- scripsit. Ofversigt af Kongliga Vetenskaps-Akad- emiens Forhandlingar 27: 243 - 358." type="journal article" year="1870">Fåhraeus 1870: 259</bibRefCitation>
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.
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</paragraph>
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<treatmentCitationGroup id="AB607B7DFFF74A2AFD06FA67F0F966A3" pageId="14" pageNumber="550">
= 
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<taxonomicName id="4C7027D0FFF74A2AFD6EFA67F7096685" authority="Fahraeus 1870: 258" authorityName="Fahraeus" authorityPageNumber="258" authorityYear="1870" box="[709,1085,1481,1503]" class="Insecta" family="Tenebrionidae" genus="Aptila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="550" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B9048041FFF74A2AFD6EFA67F0416685" box="[709,885,1482,1503]" italics="true" pageId="14" pageNumber="550">Aptila tuberculata</emphasis>
<bibRefCitation id="EFE121A2FFF74A2AFCD6FA64F7096685" author="Fahraeus, O. I." box="[893,1085,1481,1503]" pageId="14" pageNumber="550" pagination="243 - 358" refId="ref25701" refString="Fahraeus, O. I. 1870. Coleoptera Caffrariae, annis 1838 - 1845 a J. A. Wahlberg collecta. Heteromera de- scripsit. Ofversigt af Kongliga Vetenskaps-Akad- emiens Forhandlingar 27: 243 - 358." type="journal article" year="1870">Fåhraeus 1870: 258</bibRefCitation>
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[syn. by 
<treatmentCitation id="0AD17A42FFF74A2AFD6EFA49F0F666A3" author="Haag-Rutenberg, G." box="[709,962,1508,1530]" page="32" pageId="14" pageNumber="550" year="1871">
<bibRefCitation id="EFE121A2FFF74A2AFD6EFA49F0F666A3" author="Haag-Rutenberg, G." box="[709,962,1508,1530]" pageId="14" pageNumber="550" pagination="21 - 111" refId="ref27168" refString="Haag-Rutenberg, G. 1871. Beitrage zur Familie der Tenebrioniden (II. Stuck). Coleopterologische Hefte 7: 21 - 111." type="journal article" year="1871">Haag-Rutenberg 1871: 32</bibRefCitation>
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].
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<treatmentCitationGroup id="AB607B7DFFF74A2AFD06FA52F0E76575" pageId="14" pageNumber="550">
<emphasis id="B9048041FFF74A2AFD06FA52F18F654E" bold="true" box="[685,699,1535,1556]" pageId="14" pageNumber="550">=</emphasis>
<treatmentCitation id="0AD17A42FFF74A2AFD6EF9ADF7A7654E" author="Peringuey, L. A." box="[709,1171,1535,1557]" page="308" pageId="14" pageNumber="550" year="1899">
<taxonomicName id="4C7027D0FFF74A2AFD6EF9ADF7A7654E" authority="Peringuey 1899: 308" authorityName="Peringuey" authorityPageNumber="308" authorityYear="1899" box="[709,1171,1535,1557]" class="Insecta" family="Tenebrionidae" genus="Micrantereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="550" phylum="Arthropoda" rank="species" species="litigiosus">
<emphasis id="B9048041FFF74A2AFD6EF9ADF098654E" box="[709,940,1535,1557]" italics="true" pageId="14" pageNumber="550">Micrantereus litigiosus</emphasis>
<bibRefCitation id="EFE121A2FFF74A2AFC11FA52F7A7654E" author="Peringuey, L. A." box="[954,1171,1535,1557]" pageId="14" pageNumber="550" pagination="240 - 330" refId="ref28440" refString="Peringuey, L. A. 1899. Fifth contribution to the South-African coleopterous fauna. Annals of the South African Museum 1: 240 - 330." type="journal article" year="1899">Péringuey 1899: 308</bibRefCitation>
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[syn. by 
<treatmentCitation id="0AD17A42FFF74A2AFCB2F9B7F0FE6575" author="Gebien, H." box="[793,970,1561,1584]" page="905" pageId="14" pageNumber="550" year="1943">
<bibRefCitation id="EFE121A2FFF74A2AFCB2F9B7F0FE6575" author="Gebien, H." box="[793,970,1561,1584]" pageId="14" pageNumber="550" pagination="839 - 926" refId="ref26746" refString="Gebien, H. 1943. Katalog der Tenebrioniden (Col. Heteromera). Teil 3. Mitteilungen der Munchener Entomologischen Gesellschaft 33: 839 - 926 (810 - 841)." type="journal article" year="1943">Gebien 1943: 905</bibRefCitation>
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].
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<paragraph id="8BCF5C53FFF74A2AFD6EF9E9F1CA65CA" blockId="14.[685,1171,1604,1733]" pageId="14" pageNumber="550">
<emphasis id="B9048041FFF74A2AFD6EF9E9F0036503" bold="true" box="[709,823,1604,1625]" pageId="14" pageNumber="550">Type Data.</emphasis>
<taxonomicName id="4C7027D0FFF74A2AFC96F9E9F7536503" authority="Fahraeus, 1870" authorityName="Fahraeus" authorityYear="1870" box="[829,1127,1604,1626]" class="Insecta" family="Tenebrionidae" genus="Aptila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="550" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="B9048041FFF74A2AFC96F9E9F0FC6500" box="[829,968,1604,1626]" italics="true" pageId="14" pageNumber="550">Aptila costata</emphasis>
<bibRefCitation id="EFE121A2FFF74A2AFC66F9E9F7536503" author="Fahraeus, O. I." box="[973,1127,1604,1626]" pageId="14" pageNumber="550" pagination="243 - 358" refId="ref25701" refString="Fahraeus, O. I. 1870. Coleoptera Caffrariae, annis 1838 - 1845 a J. A. Wahlberg collecta. Heteromera de- scripsit. Ofversigt af Kongliga Vetenskaps-Akad- emiens Forhandlingar 27: 243 - 358." type="journal article" year="1870">Fåhraeus, 1870</bibRefCitation>
</taxonomicName>
, 
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<typeStatus id="54CBE2F1FFF74A2AFBDBF9E9F1DA652E" pageId="14" pageNumber="550" type="holotype">holotype</typeStatus>
</emphasis>
, unsexed (Naturhistoriska riksmuseet): no data.
</paragraph>
<paragraph id="8BCF5C53FFF74A2AFD6EF939F77A659F" blockId="14.[685,1171,1604,1733]" pageId="14" pageNumber="550">
<taxonomicName id="4C7027D0FFF74A2AFD6EF939F73865F3" authority="Fahraeus, 1870" authorityName="Fahraeus" authorityYear="1870" box="[709,1036,1684,1706]" class="Insecta" family="Tenebrionidae" genus="Aptila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="550" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B9048041FFF74A2AFD6EF939F04765F3" box="[709,883,1684,1705]" italics="true" pageId="14" pageNumber="550">Aptila tuberculata</emphasis>
<bibRefCitation id="EFE121A2FFF74A2AFCD3F939F73865F3" author="Fahraeus, O. I." box="[888,1036,1684,1706]" pageId="14" pageNumber="550" pagination="243 - 358" refId="ref25701" refString="Fahraeus, O. I. 1870. Coleoptera Caffrariae, annis 1838 - 1845 a J. A. Wahlberg collecta. Heteromera de- scripsit. Ofversigt af Kongliga Vetenskaps-Akad- emiens Forhandlingar 27: 243 - 358." type="journal article" year="1870">Fåhraeus, 1870</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B9048041FFF74A2AFBBEF939F75965F3" bold="true" box="[1045,1133,1684,1705]" pageId="14" pageNumber="550">
<typeStatus id="54CBE2F1FFF74A2AFBBEF939F75965F3" box="[1045,1133,1684,1705]" pageId="14" pageNumber="550" type="holotype">holotype</typeStatus>
</emphasis>
, female (Naturhistoriska riksmuseet): no data.
</paragraph>
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<taxonomicName id="4C7027D0FFF64A2BFF6EFF6FF165638D" authority="Peringuey, 1899" authorityName="Peringuey" authorityYear="1899" box="[197,593,194,216]" class="Insecta" family="Tenebrionidae" genus="Micrantereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="551" phylum="Arthropoda" rank="species" species="litigiosus">
<emphasis id="B9048041FFF64A2BFF6EFF6FF293638D" box="[197,423,194,215]" italics="true" pageId="15" pageNumber="551">Micrantereus litigiosus</emphasis>
<materialsCitation id="3B18560EFFF64A2BFE1BFF6FF165638D" box="[432,593,194,216]" collectorName="Miss M. Good" country="Botswana" location="Bechuanaland" pageId="15" pageNumber="551" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFE121A2FFF64A2BFE1BFF6FF165638D" author="Peringuey, L. A." box="[432,593,194,216]" pageId="15" pageNumber="551" pagination="240 - 330" refId="ref28440" refString="Peringuey, L. A. 1899. Fifth contribution to the South-African coleopterous fauna. Annals of the South African Museum 1: 240 - 330." type="journal article" year="1899">Péringuey, 1899</bibRefCitation>
</materialsCitation>
</taxonomicName>
, 
<emphasis id="B9048041FFF64A2BFDF4FF6FF3ED63AB" bold="true" pageId="15" pageNumber="551">
<typeStatus id="54CBE2F1FFF64A2BFDF4FF6FF3ED63AB" pageId="15" pageNumber="551" type="holotype">holotype</typeStatus>
</emphasis>
, unsexed (Cape Museum): “ 
<collectingCountry id="F3671CC3FFF64A2BFDA3FF71F1A663A8" box="[520,658,220,242]" name="Botswana" pageId="15" pageNumber="551">Bechuanaland</collectingCountry>
(Ramoutsa). 
<collectorName id="26853985FFF64A2BFE82FF5AF28D6257" box="[297,441,247,269]" pageId="15" pageNumber="551">Miss M. Good</collectorName>
”.
</paragraph>
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<paragraph id="8BCF5C53FFF64A2BFF6EFEBCF3D8621B" blockId="15.[173,659,194,321]" pageId="15" pageNumber="551">
<emphasis id="B9048041FFF64A2BFF6EFEBCF276627D" bold="true" box="[197,322,273,295]" pageId="15" pageNumber="551">Distribution.</emphasis>
<collectingCountry id="F3671CC3FFF64A2BFEEEFEBFF2FD627D" box="[325,457,274,295]" name="Botswana" pageId="15" pageNumber="551">BOTSWANA</collectingCountry>
: Ramoutsa (
<bibRefCitation id="EFE121A2FFF64A2BFD9EFEBFF3D5621B" author="Peringuey, L. A." pageId="15" pageNumber="551" pagination="240 - 330" refId="ref28440" refString="Peringuey, L. A. 1899. Fifth contribution to the South-African coleopterous fauna. Annals of the South African Museum 1: 240 - 330." type="journal article" year="1899">Péringuey 1899</bibRefCitation>
).
</paragraph>
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