Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet Author Yang, Qi-cheng 9CB67D26-5876-4E98-BEB4-B90C61F7BD88&9EF6D28F-C1F7-44EC-9331-386E8CAD95F9 Hubei Insect Resources Utilization and Sustainable Pest Management Key Laboratory, College of Plant Science & Technology of Huazhong Agriculture University, Wuhan, Hubei, 430070, China. Plant Protection College, Shenyang Agricultural University, Shenyang, Liaoning, 110866, China. Institute of Plateau Ecology, Tibet Agriculture & Animal Husbandry University, Linzhi, Xizang, P. R. 860000, China. China Agricultural University, Beijing, 100193, China. qichengyang@foxmail.com&liyan_syau@163.com Author Pan, Zhao-hui 7A386EA0-4805-4697-9225-091FEE032D13 panzhaohui2005@163.com Author Liu, Xiao-yan 40D23499-5993-4067-BF84-22907C5C8B47 Hubei Insect Resources Utilization and Sustainable Pest Management Key Laboratory, College of Plant Science & Technology of Huazhong Agriculture University, Wuhan, Hubei, 430070, China. Plant Protection College, Shenyang Agricultural University, Shenyang, Liaoning, 110866, China. Institute of Plateau Ecology, Tibet Agriculture & Animal Husbandry University, Linzhi, Xizang, P. R. 860000, China. China Agricultural University, Beijing, 100193, China. liuxiaoyan@mail.hzau.edu.cn;yangding@cau.edu.cn Author Yang, Ding FD9077E0-D8D5-4A3A-80FD-2862726AA066 liuxiaoyan@mail.hzau.edu.cn;yangding@cau.edu.cn text European Journal of Taxonomy 2021 2021-08-27 764 1 152 192 http://dx.doi.org/10.5852/ejt.2021.764.1477 journal article 4095 10.5852/ejt.2021.764.1477 e9baa250-ec84-440f-96bf-06b55f6738e3 2118-9773 5514464 8B5799F9-487D-4A9F-BF9F-54ADA06056FF Tipula ( Vestiplex ) nayogabuensis sp. nov. urn:lsid:zoobank.org:act: D91DF543-8018-4FD5-8E99-DCB2AAF3E8FF Figs 21–29 , 86 Diagnosis Vertex brownish black, relatively protruded. Prescutum virtually brownish black, black stripes distinct, all stripes with dark black margins. Scutum virtually brownish black. Scutellum dark yellow with brownish black mid-longitudinal stripe. Mediotergite reddish dark brown, dried with pale gray pruinescence and black mid-longitudinally. Thoracic pleuron dark brown with pale gray pruinescence. Posterior margin of tergite 9 with long setae, pair of weak arms arising from ventral surface of tergite 9, slender hidden bridge between them. Etymology The species is named after the type locality Nayogabu Snow Mountain. Material examined Holotype CHINA • ♂ ; Tibet , Bomi , Yigong ; 30°16′19″ N , 94°48′28″ E ; 2290 m a.s.l. ; 7 May 2017 ; Qicheng Yang leg.; light trap ; CAU . Paratypes CHINA • 8 ♂♂ ; same collection data as for holotype; CAU . Description Male MEASUREMENTS. Body length 16.5–18.5 mm, wing length 20–22.0 mm, antenna length 4.5 mm (n = 9). HEAD ( Figs 21, 23 ). Mainly brownish yellow. Occipital mark dark brown, vertex brownish black, relatively protruded. Rostrum and nasus dark yellow, nasus relatively long. Setae on head dark brown. Scape dark yellow, pedicel pale yellow, flagellum brownish black except first flagellar segment yellowish brown. Proboscis and palpus yellowish gray. THORAX ( Figs 21–23 ). Mainly dark brown, heavily pale gray pruinescence. Pronotum brown with yellow setae. Prescutum virtually brownish black, black stripes distinct, all stripes with dark black margin. Scutum virtually brownish black. Scutellum dark yellow with brownish black mid-longitudinal stripe. Mediotergite reddish dark brown, dried with pale gray pruinescence and black mid-longitudinally. Thoracic pleuron dark brown with pale gray pruinescence. Parascutellum dark brown. Meron brown. Setae on thorax pale yellow. Legs with coxae and femora yellow, femora with brownish black apexes; tibiae and tarsi dark brown. Setae on legs dark brown. Wing variegated brown; pterostigma brown; petiole of cell m distinctly shorter than discal cell ( Fig. 24 ). Halter with stem yellow, knob dark brown. ABDOMEN ( Figs 21, 23 ). Mainly yellow. Abdominal tergites each with three stripes, middle stripe of tergites 1–2 brownish black, wide, others brown; bilateral stripes black. Abdominal segments 6–9 dark black. Setae on abdomen yellow. Figs 21–24. Tipula ( Vestiplex ) nayogabuensis sp. nov. , holotype, ♂ (CAU). 21 . Habitus, lateral view. 22 . Parasitic mites. 23 . Habitus, dorsal view. 24 . Wing. Abbreviations: see Material and methods. Scale bars = 1.0 mm. Figs 25–29. Tipula ( Vestiplex ) nayogabuensis sp. nov. , holotype, ♂ (CAU). 25 . Hypopygium, lateral view. 26 . Outer gonostylus, lateral external view. 27 . Inner gonostylus, lateral external view. 28 . Tergite nine, dorsal view. 29 . Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 25, 28–29 = 0.5 mm; 26–27 = 0.1 mm. HYPOPYGIUM ( Figs 25–29 ). Gonocoxite hardly elongated, unseparated from sternite 9. Tergite 9 longitudinally divided into two parts, posterior margin with long setae, a pair of weak arms arising from ventral surface of tergite 9, a slender hidden bridge between them ( Fig. 28 ). Appendages of sternite 9 slender ( Fig. 29 ). Apex and middle of outer gonostylus slightly wide ( Fig. 26 ). Inner gonostylus with concavity at middle, posterior part with many bristles, lower margin with bristled raise; base of lower beak inward-rolled, enwrapped black lower beak; upper beak yellow, lower margin black, base with a slight raise ( Fig. 27 ). Apex of adminiculum hooked. Female Unknown or unrecognized. Distribution China : Tibet . Remarks The new species is similar to T . ( V .) zayulensis from Chayu (= Zayu) of Tibet , but the latter differs in the following characters: prescutum and scutum yellowish brown; thoracic pleuron brownish yellow; all sterno-pleurite with dark brown margin. The outer gonostylus is different from that of T . ( V .) nayogabuensis sp. nov.