Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula Author Mertlik, Josef Pohřebačka 34, CZ- 53345 Opatovice nad Labem, Czech Republic mertlik@elateridae.com Author Németh, Tamás Hungarian Natural History Museum, Department of Zoology, Baross u. 13, H- 1088 Budapest, Hungary haesito@gmail.com Author Kundrata, Robin Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, CZ- 77146 Olomouc, Czech Republic robin.kundrata@upol.cz text Zootaxa 2017 2017-01-13 4220 1 1 63 journal article 7453 10.11646/zootaxa.4220.1.1 eb597d59-1f6e-412d-a99a-de1a67e7cc94 1175-5326 4670787 D74BC90C-84CC-4788-9048-54F5C8521B32 Dima bruhai sp. nov. Figs 10–13 , 121 , 163 , 178 , 223 , 244 . Type material. Holotype : GREECE : distr. Kavála , Pangéo Mts. , 2 km S of Paleochori , beechwood, 1080 m ( 40°55'20.25"N , 24°10'45.56"E ), 3.VI.2014 , ♂ , J. Mertlik leg. ( PCJM ) . Paratypes : GREECE : distr. Kavála , Mt. Pangeo , 1050 m (ca. 40°55'14.15"N , 24°10'30.76"E ), 5.VI.2006 , 4 ♂♂ , 2 ♀♀ , beechwood, in trap , A. Müller leg. ( PCME ) ; Pangeo Mts. , 1375 m , 15.VI.2007 , 1 ♂ , 1 ♀ , S. Peslier leg. ( PCGP ) ; distr. Kavála , Pangéo Mts. , 2 km S of Paleochori , beechwood, 1080 m ( 40°55'20.25"N , 24°10'45.56"E ), 3.VI.2014 , 27 ♂♂ , 10 ♀♀ , J. Mertlik leg. ( PCJM ) ; distr. Kavála , Pangéo Mts. , 2.5 km S of Paleochori , beechwood ( 40°55'13.79"N , 24°11'1.79"E ), 3.VI.2014 , 3 ♂♂ , 2 ♀♀ , J. Mertlik leg. ( PCJM ) ; dtto, 3 ♂♂ (PCRK); distr. Kavála , Pangéo Mts. , 2 km S of Paleochori , beechwood, 1080 m ( 40°55'20.25"N , 24°10'45.56"E ), 6.VI.2015 , 28 ♂♂ , 11 ♀♀ , J. Mertlik leg. ( PCJM ) ; dtto, 33 ♂♂, 4 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 20 ♂♂, 11 ♀♀, P. Brůha leg. (PCPB); dtto, 6 ♂♂, 2 ♀♀, T. Németh leg. (HNHM); dtto, 14 ♂♂, 6 ♀♀ (PCRK); 1.–10.VII.2015 , 1 ♂, 4 ♀♀, Manfred Egger leg. (PCME). Diagnosis. Dima bruhai sp. nov. is a larger species characterized by having the pubescence of the pronotal sides semi-erect along the whole length, gradually longer towards posterior angles, and moderately matt elytra with dense, moderately long pubescence ( Figs 10–13 , 121 ). The other Dima spp. in the surrounding area differ from D. bruhai sp. nov. by different pubescence of pronotum and elytra. Dima schimmeli Platia & Gudenzi, 2009 has body surface covered with the long, straight, semi-erect pubescence ( Figs 102–104 , 156 ). Dima orientalis sp. nov. has pubescence of the pronotal sides almost decumbent, with longer semi-erect hairs only at hind angles, and elytral pubescence much sparser ( Figs 77–79 , 148 ). Dima kozufensis sp. nov. has the more compact body shape, pubescence of the pronotal sides decumbent anteriorly, pubescence of elytra slightly sparser and shorter, scutellum basally emarginate and more coarsely punctate, and apical lobe of paramera slightly shorter ( Figs 50–52 , 138 , 191 ). Dima lebenbaueri Schimmel & Platia, 2008 has less matt body surface, longer antennomeres II and III, longer pronotal pubescence, and sparser elytral pubescence (much sparser in the larger specimens from the type locality; Figs 53–56 , 139–140 ). Description. Holotype , male. Body rather large, 14.8 mm long, 5.9 mm wide, matt. Body dark brown, head paler, antennae (except darker antennomeres II–III), pronotal angles, legs, elytral sutura and margins red-brown; pubescence yellowish ( Fig. 10 ). Head including eyes 0.5 times as wide as pronotum, frons with shallow v-shaped depression in the middle; punctation moderately dense and coarse, intervals between punctures shiny posterad, rugose anterad; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Antennae surpassing the posterior angles of pronotum of about four segments; antennomeres II–III short, antennomere II about 1.5 times longer than wide, antennomere III about 1.3 times longer than wide, length ratio of antennomeres II–IV 1.2: 1.0: 2.0, antennomeres IV–X gradually widened apically, less than 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae. Pronotum 1.4 times wider than long, widest behind middle, moderately convex dorsally; sides evenly convex, straight in lateral view; anterior angles obtuse; posterior angles prominent, sharp, distinctly produced outward; punctation dense, coarse, moderately deep, punctures on disc equally distributed, of about the same size; intervals between punctures about two times a diameter of puncture, moderately shiny. Pubescence on pronotal disc short, decumbent to semi-erect; that on pronotal sides semi-erect along the whole length, gradually longer towards posterior angles ( Fig. 121 ). Prosternum finely, densely punctate, with decumbent to semi-ercet setae, denser at margins; prosternal lobe coarsely punctate, with longer pubescence. Scutellum sub-pentagonal, wide, flattened, sides sub-parallel, in lateral view under the plane of elytra ( Fig. 163 ), frontal margin widely rounded, gradually declined; apex rounded, punctation coarse, dense; pubescence fine, moderately long, semi-erect. Elytra sub-parallel, 3.3 times longer than pronotum, moderately convex, widest at posterior two thirds; striae well developed along the whole elytral length, irregularly interrupted by coarse sparse punctures; surface of interstices flattened, moderately matt, densely and coarsely punctate; punctures distinct, equally distributed, suboval; pubescence dense, moderately long. Aedeagus elongate; paramera long, narrow, its apical lobe long, narrow, subapical tooth minute, subacute ( Fig. 178 ). Female ( Fig. 11 ). Like male but with more robust, suboval body, antennae shorter (about 2.5 segments surpassing posterior angles of pronotum), with segments IV–X shorter (about 1.7 times longer than wide), pronotum slightly wider and more convex, widest at middle. The sclerotized spines of bursa copulatrix as in Fig. 223 . Intraspecific variability. Body length: 11.8–17.1 mm . Some specimens are paler and have more elongate body shape (2.6 times longer than wide) with longer pronotum, including the largest known specimen ( 17.1 mm ), which is also slightly asymmetrical ( Figs 10–13 ). Additionally, some specimens have antennomeres I–III or II–III differently colored than remaining antennomeres (either paler or darker, the latter situation is present in the holotype ). Distribution. Greece (Pangeo Mts; Fig. 244 ). Etymology. The specific epithet is a patronym in honor of Mr. Petr Brůha ( Ústí nad Labem , Czech Republic ), who collected a part of the type series.