<document id="1498528161F9C2C9FEF4EA99484BD6FA" ID-DOI="10.5281/zenodo.13203792" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203792" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722658458551" checkinUser="felipe" docAuthor="Minkina, Ÿukasz, Anichtchenko, Alexander, Vasiljeva, Anastasija &amp; Skelley, Paul E." docDate="2022" docId="03BD44335E5BFFE5FCD826D800F2FF68" docLanguage="en" docName="BalticJColeopterol.22.1.81-101.pdf" docOrigin="Baltic Journal of Coleopterology 22 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.4:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="4" docTitle="Rhyparus paraphilippinensis Ochi &amp; Kakizoe 2021" docType="treatment" docVersion="3" lastPageNumber="92" masterDocId="FF843C4B5E51FFEEFF8A22690417FFB0" masterDocTitle="Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females" masterLastPageNumber="101" masterPageNumber="81" pageNumber="91" updateTime="1722698086404" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="142019366167C7858A7658AA28503BC3">Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females</mods:title>
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<mods:namePart id="356E4DCEB3D1C441BB95974D8BF460D4">Minkina, Ÿukasz</mods:namePart>
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<mods:namePart id="FA90FBD707D602D1153D9BA716CBF15E">Anichtchenko, Alexander</mods:namePart>
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<mods:namePart id="5AB65853C0939A39365DA3F0ECB8E610">Vasiljeva, Anastasija</mods:namePart>
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<mods:namePart id="90EFA5313AD264E9C122EDA2BE3CF531">Skelley, Paul E.</mods:namePart>
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<mods:title id="AFCD6E0B0FFDDACE6C258AA69F1F54C0">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="736E7C2E5FF6B95AD65FB02353832049">2022</mods:date>
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<taxonomicName id="4C148EA65E5BFFE4FCD826D8079FFB59" ID-CoL="8TJR4" authority="Ochi &amp; Kakizoe, 2021" authorityName="Ochi &amp; Kakizoe" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="paraphilippinensis">
<emphasis id="B96029375E5BFFE4FCD826D80084FB79" bold="true" box="[850,1171,1201,1225]" italics="true" pageId="10" pageNumber="91">Rhyparus paraphilippinensis</emphasis>
<bibRefCitation id="EF8588D45E5BFFE4FB1026DB079FFB59" author="Ochi, T. &amp; Kakizoe, S. &amp; Minkina, Y. &amp; Kawahara, M. &amp; Cabras, A. A." pageId="10" pageNumber="91" pagination="17 - 42" refId="ref9157" refString="Ochi, T., Kakizoe, S., Minkina, Y., Kawahara, M. &amp; Cabras, A. A. 2021: A revisional study of the genus Rhyparus from the Philippines. Kogane, 24: 17 - 42." type="journal article" year="2021">Ochi &amp; Kakizoe, 2021</bibRefCitation>
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(
<figureCitation id="132FE9A05E5BFFE4FCD0269A07CAFABA" box="[858,989,1267,1290]" captionStart="Figs" captionStartId="11.[227,277,1693,1716]" captionTargetBox="[367,1150,758,1657]" captionTargetId="figure-16@11.[366,1153,752,1660]" captionTargetPageId="11" captionText="Figs. 13–15. Rhyparus paraphilippinensis Ochi &amp; Kakizoe, 2021.,: 13 – dorsal view; 14 – ventral view; 15 – lateral view. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203804" httpUri="https://zenodo.org/record/13203804/files/figure.png" pageId="10" pageNumber="91">Figs. 13–15</figureCitation>
, 
<figureCitation id="132FE9A05E5BFFE4FC63269A0011FABA" box="[1001,1030,1267,1290]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 20–25. Heads: 20 – R. bacanensis sp. nov.,, holotype; 21 – R. buruensis sp. nov.,, holotype; 22 – R. fijiensis sp. nov.,, holotype; 23 – R. stebnickae sp. nov.,, holotype; 24 – R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 25 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203808" httpUri="https://zenodo.org/record/13203808/files/figure.png" pageId="10" pageNumber="91">24</figureCitation>
, 
<figureCitation id="132FE9A05E5BFFE4FB9B269A003AFABA" box="[1041,1069,1267,1290]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 26–33.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 – R. bacanensis sp. nov.,, holotype; 27 – R. buruensis sp. nov.,, holotype; 28 – R. fijiensis sp. nov.,, holotype; 29 – R. stebnickae sp. nov.,, holotype; 30 – R. approximans Fairmaire, 1893,; 31 – R. loebli Paulian, 1983,; 32 – R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13203810" httpUri="https://zenodo.org/record/13203810/files/figure.png" pageId="10" pageNumber="91">32</figureCitation>
)
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<emphasis id="B96029375E5BFFE4FCD8275D009AFAFB" box="[850,1165,1332,1355]" italics="true" pageId="10" pageNumber="91">Rhyparus paraphilippinensis</emphasis>
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, figs: 5, 18, 30, 38, 45. 
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locality: “ 
<collectingCountry id="F303B5B55E5BFFE4FCD6271C07CCFA3C" box="[860,987,1397,1420]" name="Philippines" pageId="10" pageNumber="91">Philippines</collectingCountry>
, 
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, 
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, 
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”.
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<treatmentCitationGroup id="AB04D20B5E5BFFE4FCD827DF0123FA5D" pageId="10" pageNumber="91">
<taxonomicName id="4C148EA65E5BFFE4FCD827DF00B9FA7D" ID-CoL="4SV4J" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[850,1198,1462,1485]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B96029375E5BFFE4FCD827DF0001FA7D" box="[850,1046,1462,1485]" italics="true" pageId="10" pageNumber="91">Rhyparus gracilis</emphasis>
Arrow, 1905
</taxonomicName>
: 
<treatmentCitation id="0AB5D3345E5BFFE4FB3327DF0012FA5D" author="Anichtchenko, A. V. &amp; Minkina, L. &amp; Vasiljeva, A. &amp; Medina, M. N. D." httpUri="http://treatment.plazi.org/id/03AA8121533BFFDF53B66A76BAF7F7A7" page="102" pageId="10" pageNumber="91" year="2021">
<bibRefCitation id="EF8588D45E5BFFE4FB3327DF0012FA5D" author="Anichtchenko, A. V. &amp; Minkina, L. &amp; Vasiljeva, A. &amp; Medina, M. N. D." pageId="10" pageNumber="91" pagination="99 - 111" refId="ref8853" refString="Anichtchenko, A. V., Minkina, L., Vasiljeva, A. &amp; Medina, M. N. D. 2021: A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao. A cta entomologica Musei nationalis Pragae 61 (1): 99 - 111." type="journal article" year="2021">
Anichtchenko 
<emphasis id="B96029375E5BFFE4FCD827BE079EFA5E" box="[850,905,1495,1518]" italics="true" pageId="10" pageNumber="91">et al.</emphasis>
, 2021: 102
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, figs: 3a­c, 9, 15, 21, 27a­b.
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</paragraph>
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<emphasis id="B96029375E5BFFE4FCD8247E00B1F99F" bold="true" box="[850,1190,1559,1583]" pageId="10" pageNumber="91">Additional material examined.</emphasis>
(22exx.): 
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8exx.: 
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, 
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, 
<location id="8ECBA3FE5E5BFFE4FBC6245100F7F9FF" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5BFFE4FBC6245100F7F9FF" box="[1100,1248,1592,1615]" country="Philippines" county="Eureka Mt." latitude="8.691617" longLatPrecision="1" longitude="125.23785" municipality="Gingoog" name="Bagongsilang" pageId="10" pageNumber="91">Bagongsilang</location>
, 1996, leg. 
<collectorName id="26E190F35E5BFFE4FCD824300780F9C0" box="[850,919,1625,1648]" pageId="10" pageNumber="91">Bocak</collectorName>
(
<collectionCode id="ED056DE05E5BFFE4FC2F243007E0F9C0" box="[933,1015,1625,1648]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="91">NMPC</collectionCode>
); 3exx.: 
<collectingCountry id="F303B5B55E5BFFE4FBDF243000C6F9C0" box="[1109,1233,1625,1648]" name="Philippines" pageId="10" pageNumber="91">Philippines</collectingCountry>
, 
<location id="8ECBA3FE5E5BFFE4FB562430015BF9C0" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5BFFE4FB562430015BF9C0" box="[1244,1356,1625,1648]" country="Philippines" county="Eureka Mt." latitude="8.691617" longLatPrecision="1" longitude="125.23785" municipality="Gingoog" name="Mindanao" pageId="10" pageNumber="91">Mindanao</location>
, 
<collectingMunicipality id="6BCF6F5F5E5BFFE4FCD8241007A0F920" box="[850,951,1657,1680]" pageId="10" pageNumber="91">Gingoog</collectingMunicipality>
, 
<collectingCounty id="62CA8DA95E5BFFE4FC4124100045F920" box="[971,1106,1657,1680]" pageId="10" pageNumber="91">Eureka Mt.</collectingCounty>
, 
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, 
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, 
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, 
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, leg. 
<collectorName id="26E190F35E5BFFE4FB7D24F30768F961" pageId="10" pageNumber="91">A.Vasiljeva</collectorName>
(
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); 
<specimenCount id="9D123EAC5E5BFFE4FC6724D30009F961" box="[1005,1054,1722,1745]" count="1" pageId="10" pageNumber="91" type="generic">1ex.</specimenCount>
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: the same but: leg. A.Anicht­
</paragraph>
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<emphasis id="B96029375E5AFFE5FD8A22F807FFFF14" box="[512,1000,145,164]" italics="true" pageId="11" pageNumber="92">Minkina Ł., Anichtchenko A., Vasiljeva A., Skelley P.E.</emphasis>
</paragraph>
<paragraph id="8BABF5255E5AFFE5FF6922A80697FDA8" blockId="11.[227,740,193,536]" pageId="11" pageNumber="92">
<materialsCitation id="3B7CFF785E5AFFE5FF6922A805D5FE48" collectingDate="2022-03-21" collectingDateMax="2022-03-28" collectingDateMin="2022-03-21" collectionCode="DUBC" collectorName="A. Vasiljeva &amp; Apo Mt. Rng" country="Philippines" county="Mindanao" elevation="1200" latitude="7.12135" location="Claveria rd." longLatPrecision="1" longitude="125.31263" municipality="Misamis" pageId="11" pageNumber="92" specimenCount="3" stateProvince="Davao del Sur">
chenko (
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); 
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: 
<collectingCountry id="F303B5B55E5AFFE5FE6E22A80677FF68" box="[484,608,193,216]" name="Philippines" pageId="11" pageNumber="92">Philippines</collectingCountry>
, 
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, 
<collectingMunicipality id="6BCF6F5F5E5AFFE5FF6922880555FF48" box="[227,322,225,248]" pageId="11" pageNumber="92">Misamis</collectingMunicipality>
or., 
<location id="8ECBA3FE5E5AFFE5FEF122880615FF48" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FEF122880615FF48" box="[379,514,225,248]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Claveria rd." pageId="11" pageNumber="92" stateProvince="Davao del Sur">Claveria rd.</location>
, 
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</quantity>
, 8.701522, 125.014624, 
<date id="FFAAD3E55E5AFFE5FEF02368060EFEA8" box="[378,537,257,280]" pageId="11" pageNumber="92" value="2022-03-26" valueMax="2022-03-28" valueMin="2022-03-26">
<collectingDate id="EFEE2A0D5E5AFFE5FEF02368060EFEA8" box="[378,537,257,280]" pageId="11" pageNumber="92" value="2022-03-26" valueMax="2022-03-28" valueMin="2022-03-26">26­28.iii.2022</collectingDate>
</date>
, leg. 
<collectorName id="26E190F35E5AFFE5FDD5236806F6FEA8" box="[607,737,257,280]" pageId="11" pageNumber="92">A.Vasiljeva</collectorName>
(
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); 
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: 
<collectingCountry id="F303B5B55E5AFFE5FE0B234805EEFE88" box="[385,505,289,312]" name="Philippines" pageId="11" pageNumber="92">Philippines</collectingCountry>
, 
<location id="8ECBA3FE5E5AFFE5FD8823480678FE88" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FD8823480678FE88" box="[514,623,289,312]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mindanao" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mindanao</location>
, 
<collectingRegion id="49D03BC75E5AFFE5FDF223480510FEE8" country="Philippines" name="Davao del Sur" pageId="11" pageNumber="92">Davao del Sur</collectingRegion>
, 
<collectorName id="26E190F35E5AFFE5FE98232805B7FEE8" box="[274,416,321,344]" pageId="11" pageNumber="92">
<location id="8ECBA3FE5E5AFFE5FE982328057BFEE8" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FE982328057BFEE8" box="[274,364,321,344]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Apo Mt." pageId="11" pageNumber="92" stateProvince="Davao del Sur">
Apo 
<location id="8ECBA3FE5E5AFFE5FECF2328057BFEE8" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FECF2328057BFEE8" box="[325,364,321,344]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mt. Rng" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mt.</location>
</location>
Rng
</collectorName>
, 
<location id="8ECBA3FE5E5AFFE5FE2023280602FEE8" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FE2023280602FEE8" box="[426,533,321,344]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mabanlas" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mabanlas</location>
, 
<quantity id="4CEC58C05E5AFFE5FD942328067CFEE8" box="[542,619,321,344]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="92" unit="m" value="1200.0">
<elevation id="003912165E5AFFE5FD942328067CFEE8" box="[542,619,321,344]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="92" unit="m" value="1200.0">1200m</elevation>
</quantity>
, 7.120251, 125.317203, 
<date id="FFAAD3E55E5AFFE5FEFD23080603FEC8" box="[375,532,353,376]" pageId="11" pageNumber="92" value="2022-03-23" valueMax="2022-03-24" valueMin="2022-03-23">
<collectingDate id="EFEE2A0D5E5AFFE5FEFD23080603FEC8" box="[375,532,353,376]" pageId="11" pageNumber="92" value="2022-03-23" valueMax="2022-03-24" valueMin="2022-03-23">23­24.iii.2022</collectingDate>
</date>
, leg. 
<collectorName id="26E190F35E5AFFE5FDDD230806F6FEC8" box="[599,737,353,376]" pageId="11" pageNumber="92">A. Vasiljeva</collectorName>
(
<collectionCode id="ED056DE05E5AFFE5FF6023E8052CFE28" box="[234,315,385,408]" pageId="11" pageNumber="92">DUBC</collectionCode>
); 
<specimenCount id="9D123EAC5E5AFFE5FEC423E80568FE28" box="[334,383,385,408]" count="1" pageId="11" pageNumber="92" type="generic">1ex.</specimenCount>
: the same but: 
<collectingCountry id="F303B5B55E5AFFE5FDA923E80688FE28" box="[547,671,385,408]" name="Philippines" pageId="11" pageNumber="92">Philippines</collectingCountry>
, 
<location id="8ECBA3FE5E5AFFE5FD2123E80531FE08" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FD2123E80531FE08" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mindanao" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mindanao</location>
, 
<collectingRegion id="49D03BC75E5AFFE5FEBA23C805DEFE08" box="[304,457,417,440]" country="Philippines" name="Davao del Sur" pageId="11" pageNumber="92">Davao del Sur</collectingRegion>
, 
<collectorName id="26E190F35E5AFFE5FE5F23C80671FE08" box="[469,614,417,440]" pageId="11" pageNumber="92">
<location id="8ECBA3FE5E5AFFE5FE5F23C80626FE08" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FE5F23C80626FE08" box="[469,561,417,440]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Apo Mt." pageId="11" pageNumber="92" stateProvince="Davao del Sur">
Apo 
<location id="8ECBA3FE5E5AFFE5FD8023C80626FE08" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FD8023C80626FE08" box="[522,561,417,440]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mt. Rng" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mt.</location>
</location>
Rng
</collectorName>
, 
<location id="8ECBA3FE5E5AFFE5FDFB23C806CBFE08" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FDFB23C806CBFE08" box="[625,732,417,440]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mabanlas" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mabanlas</location>
, 
<quantity id="4CEC58C05E5AFFE5FF6923A80523FE68" box="[227,308,449,472]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="92" unit="m" value="1500.0">
<elevation id="003912165E5AFFE5FF6923A80523FE68" box="[227,308,449,472]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="92" unit="m" value="1500.0">1500m</elevation>
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, 
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, 
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, 
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<collectingDate id="EFEE2A0D5E5AFFE5FD3023A8054FFE48" pageId="11" pageNumber="92" value="2022-03-21" valueMax="2022-03-23" valueMin="2022-03-21">21­ 23.iii.2022</collectingDate>
</date>
(
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)
</materialsCitation>
; 
<materialsCitation id="3B7CFF785E5AFFE5FE5A2388066AFDA8" collectingDate="2022-03-21" collectingDateMax="2022-03-24" collectingDateMin="2022-03-21" collectionCode="DUBC" collectorName="A. Vasiljeva" country="Philippines" pageId="11" pageNumber="92" specimenCount="1" stateProvince="Davao del Sur">
7exx.: the same but: 
<date id="FFAAD3E55E5AFFE5FD372388054CFDA8" pageId="11" pageNumber="92" value="2022-03-21" valueMax="2022-03-24" valueMin="2022-03-21">
<collectingDate id="EFEE2A0D5E5AFFE5FD372388054CFDA8" pageId="11" pageNumber="92" value="2022-03-21" valueMax="2022-03-24" valueMin="2022-03-21">21­ 24.iii.2022</collectingDate>
</date>
, leg. 
<collectorName id="26E190F35E5AFFE5FE1820680601FDA8" box="[402,534,513,536]" pageId="11" pageNumber="92">A. Vasiljeva</collectorName>
(
<collectionCode id="ED056DE05E5AFFE5FDAE20680662FDA8" box="[548,629,513,536]" pageId="11" pageNumber="92">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E5AFFE5FF69202905FFFD08" pageId="11" pageNumber="92" type="discussion">
<paragraph id="8BABF5255E5AFFE5FF69202905FFFD08" blockId="11.[227,737,576,696]" pageId="11" pageNumber="92">
<emphasis id="B96029375E5AFFE5FF692029054BFDE8" bold="true" box="[227,348,576,600]" pageId="11" pageNumber="92">Comment.</emphasis>
Specimens from paper of 
<bibRefCitation id="EF8588D45E5AFFE5FD07202805A1FDC8" author="Anichtchenko, A. V. &amp; Minkina, L. &amp; Vasiljeva, A. &amp; Medina, M. N. D." pageId="11" pageNumber="92" pagination="99 - 111" refId="ref8853" refString="Anichtchenko, A. V., Minkina, L., Vasiljeva, A. &amp; Medina, M. N. D. 2021: A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao. A cta entomologica Musei nationalis Pragae 61 (1): 99 - 111." type="journal article" year="2021">
Anichtchenko 
<emphasis id="B96029375E5AFFE5FEB320080566FDC8" box="[313,369,609,632]" italics="true" pageId="11" pageNumber="92">et al.</emphasis>
, 2021
</bibRefCitation>
identified by authors as 
<taxonomicName id="4C148EA65E5AFFE5FD4020080521FD28" authorityName="Arrow" authorityYear="1905" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="92" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B96029375E5AFFE5FD4020080521FD28" italics="true" pageId="11" pageNumber="92">R. gracilis</emphasis>
</taxonomicName>
indeed are 
<taxonomicName id="4C148EA65E5AFFE5FE2520E80680FD28" authorityName="Ochi &amp; Kakizoe" authorityYear="2021" box="[431,663,641,664]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="92" phylum="Arthropoda" rank="species" species="paraphilippinensis">
<emphasis id="B96029375E5AFFE5FE2520E80680FD28" box="[431,663,641,664]" italics="true" pageId="11" pageNumber="92">R. paraphilippinensis</emphasis>
</taxonomicName>
. Additionally, see: discussion.
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E5AFFE5FC8C22A900F2FF68" box="[774,1253,192,216]" pageId="11" pageNumber="92" type="distribution">
<paragraph id="8BABF5255E5AFFE5FC8C22A900F2FF68" blockId="11.[774,1253,192,216]" box="[774,1253,192,216]" pageId="11" pageNumber="92">
<emphasis id="B96029375E5AFFE5FC8C22A9078CFF68" bold="true" box="[774,923,192,216]" pageId="11" pageNumber="92">Distribution.</emphasis>
<collectingCountry id="F303B5B55E5AFFE5FC2B22A80008FF68" box="[929,1055,193,216]" name="Philippines" pageId="11" pageNumber="92">Philippines</collectingCountry>
: Mindanao Island.
</paragraph>
</subSubSection>
</treatment>
</document>