<strong> Revision of the genus <em> Aspicera </ em> Dahlbom, 1842 (Hym.: Figitidae: Aspicerinae) </ strong> Author ROS-FARRÉ, P. Author PUJADE-VILLAR, J. text Zootaxa 2013 2013-01-15 3606 1 1 110 http://dx.doi.org/10.11646/zootaxa.3606.1.1 journal article 10.11646/zootaxa.3606.1.1 1175-5326 5259487 3ED10277-CFD3-4D0D-B49A-C9F28AB6C955 Redescription of Aspicera Onychia Haliday in Walker, 1835: 162 . Type-species: Evania ediogaster Rossi, 1790 , by monotypy. Homonym of Onychia Hübner, 1816: 334 (Lepidoptera) . Aspicera Dahlbom, 1842: 6 . Type-species: Evania ediogaster Rossi, 1790 , by monotypy. Replacement name for Onychia Haliday by Reinhard, 1860: 242 . Bellona Giraud, 1860: 156 . Type-species: Evania ediogaster Rossi, 1790 , by monotypy. Unnecessary replacement name for Onychia Haliday. Homonym of Bellona Reichenbach, 1852 , (fossil bird). Heteraspidia Belizin, 1952: 299–300 . Type-species: Heteraspidia foveata Belizin, 1952 , by monotypy. Synonymyzed by Ros-Farré & Pujade-Villar, 2011a: 54 Type species: Tenthredo scutellata Villers, 1789 (= Evania ediogaster Rossi, 1790 ; see taxonomic history of Aspicera ) Head densely pubescent on lower half, glabrous to sparsely pubescent on upper half. Lateral frontal carinae conspicuous, convergent towards antennal foramina ( Figs 1b & 1e ) (in A. readae and A. elisendae they are parallel, straight with facial carinae ( Fig. 1c ). Vertex straight or slightly incised. Ocelli weakly prominent to noticeably prominent. Occipital carinae conspicuous ( Figs 32a & 44a ). Occiput carinate ( Figs 2a , 24b & 34b ) or without carinae ( Figs 7b & 8b ), usually coriaceous or alutaceous. Gena in lateral view expanded ( Fig. 44a ) or not ( Fig. 15a ) and curved ( Figs 32a & 46a ), although sometimes angled ( Fig. 27a ), more or less carinate. Frons with variable sculpture, it can be coriaceous ( Fig. 1d ), rugose or reticulate ( Figs 1c & 1f ), sometimes also punctate ( Fig. 1e ) or carinate ( Fig. 1a ). Lateral frontal carinae conspicuous, sometimes weak near lateral ocelli. Antenna filiform or subclavate; female: 13-segmented (female of A. abbreviata has 14-segmented antennae), F1 narrower at basis than apex; male: 14-segmented, F1 sharply excavated ( Fig. 2b ), with exception of A. scutellata and A. belizini , which have F1 very weakly modified ( Fig. 2a ). Mesosoma sparsely pubescent. Lateral surface of pronotum: the microsculpture is coriaceous, except in A. abbreviata , which does not have microsculpture; the macrosculpture can be transverse carinae ( Figs 43a & 44a ), rugose ( Fig. 17a ) or absent ( Figs 8a & 30a ). Pronotal plate conspicuous, subpronotal plate present, noticeably projected ahead in A. aculeata ( Fig. 1a ). Scutum coriaceous, rarely shiny ( Fig. 15b ), with ( Fig. 42b ) or without ( Fig. 27b ) transverse carinae, sometimes also punctate ( Fig. 11b ). Anteroadmedian line sharp ( Fig. 14b ) to very weak ( Fig. 8b ), reaching 1/3 to 1/2 scutum length. Median ridge sharp or effaced ( Fig. 8b ), reaching from anterior margin of scutum to median mesoscutal furrow, ( Fig. 14b ). Notauli complete, coriaceous or shiny, sometimes with transverse carinae inside. Median mesoscutal furrow reaching, at least, basal 1/3 of scutum, coriaceous or shiny, sometimes with transverse carinae inside. Parascutal sulcus anteriorly ending in front of tegula ( Figs 8a & 27a ) or continuing to anterior end of notauli ( Figs 24a & 44a ). Area between notauli and end of median mesoscutal furrow flat ( Figs 15a & 8a ) or prominent ( Figs 24a , 40a & 45a ). Mesopleuron smooth or coriaceous, with coarse transverse carinae or with a furrow on basal 1/3, with sharp sculpture below it. Scutellum with scutellar spine, which is variable in length, usually pointed ( Figs 23b & 32b ), sometimes slightly blunt ( Fig. 12b & 15b ) or near absent ( A. kerzhneri ). Scutellar disc with two large and conspicuous foveae, in lateral view flat ( Fig. 44a ) or convex ( Fig. 15a ), the end of scutellar disc can be in the same plane as scutellar spine ( Fig. 44a ) or sloping down towards apex of scutellar spine ( Figs 39a & 40a ). Propodeum pubescent, propodeal carinae sharp, dorsally parallel, ventrally divergent ( Fig. 2e ). Forewing with radial cell open, R1 present, short or long (reaching half the distance between base of R1 and wing margin), but never reaching margin of wing, R2 straight or curved, Rs+M absent or slightly impressed. Wing surface pubescence reduced or not. Marginal pubescence of wing present or absent. Metatibia always with a sharp longitudinal ridge on the posterior surface. Petiole wider than long, with conspicuous longitudinal costulae ( Fig. 2e ); 3 rd abdominal tergum saddle-shaped and smooth ( Fig. 2d ). 4 th abdominal tergum with marked punctuation, usually not reaching posterior margin of 3 rd Tergum. Diagnosis. Aspicera can be distinguished from other genera of Aspicerinae ‘ sensu stricto ’ by the following characters (Ros-Farré, 2007): - Anacharoides , Pujadella , Omalaspis and Callaspidia lack scutellar spine (present in Aspicera ). - In Balna the scutellar spine is reduced to a blunt point, and the scutellum has only one deep central fovea; in Aspicera the scutellar spine can be short but it is never reduced to a blunt point and the scutellum has two foveae. - In Paraspicera the scutellar spine is blunt, thick and wide, the scutum never lacks microsculpture and the median sulcus is always narrow; in Aspicera the scutellar spine is slender and pointed, rarely very short, but if so the scutum lacks microsculpture and is distinctly carinate transversely, and the median sulcus is wide. - In Prosaspicera there is a conspicuous median sulcus beginning at lateral ocelli and ending at occiput, with parallel longitudinal carinae at sides ( Ros-Farré & Pujade-Villar 2007 ), and the compound eyes are surrounded by a sharp continuous carina that is the union of the frontal carinae, vertex lateral carinae, occipital carina and facial impression ( Ros-Farré & Pujade-Villar 2007 ); both characters are absent in Aspicera . World key to the species of Aspicera 1. Scutellar spine almost absent, made up by a distal elongation of the circumscutellar carina ( Figs 2f , 4b )................. 2 - Scutellar spine slender and pointed, rarely very short ( Fig 13b ).................................................. 3 2. Scutellar foveae shallow, at least 1/2 scutellum length and with two longitudinal carinae inside ( Fig. 2f ). Occiput without carinae. Lateral surface of propodeum not carinate. Mesoscutum with some punctuations............ A. foveata ( Belizin, 1952 ) - Scutellar foveae well defined, about 1/3 scutellum length and with three longitudinal carinae inside ( Fig 4b ). Occiput with longitudinal carinae dorsally and transverse carinae ventrally. Lateral surface of propodeum carinate transversely. Mesoscutum with transverse carinae........................................................... A. kerzhneri ( Kovalev, 1974 ) 3. Scutellum irregularly rugose/carinate, without longitudinal carinae except for the interfoveal carina ( Fig. 5b ). Scutellar spine short and slender. Occiput rugose and carinate ( Fig. 5b ). Lateral line incomplete, ending before anterior margin of scutum ( Fig. 5a ). Wing pubescence reduced, marginal pubescence absent.................................................... 4 - Scutellum variable, but if it has rugose carinae, these are always longitudinal-like. Scutellar spine variable in length and wideness. Occiput smooth, carinate or rugose. Wing pubescence present or absent...................................... 5 4. Mesoscutum with coarse coriaceous sculpture ( Fig. 5b ). Scutellar foveae with scutellar pits ( Fig. 5b ). Female scutellar spine between 0.38 to 0.46 times length of scutellar disc...................... A. singularica Ros-Farré & Pujade-Villar n. sp. - Mesoscutum with fine coriaceous sculpture ( Fig. 6b ). Scutellar foveae without scutellar pits ( Fig. 6b ). Female scutellar spine 0.20 times length of scutellar disc................................................... A. clarimontis Kieffer, 1907 5. Scutum always clearly punctate, without transverse carinae ( Figs 7b & 11b ). Antero-admedian lines very thin or indistinct. Median ridge effaced. Scutellar spine always short, 0.3 to 0.5 times length of scutellar disc........................... 6 - Scutum sometimes punctate between notauli and lateral margins, with or without transversal carinae. Antero-admedian lines usually conspicuous. Median ridge well developed ( Figs 12b & 15b ). Scutellar spine variable in length................ 10 6. Scutellar foveae smooth, very large and wide. Scutellar disc coriaceous ( Fig. 7b )..................................................................................................... A. kovalevi Ros-Farré & Pujade-Villar n. sp. - Scutellar foveae with sharp to effaced longitudinal carinae, in A. jantonii it is nearly smooth, variable in size. Scutellar disc carinate longitudinally and emarginate ( Figs 8b & 10b ).......................................................... 7 7. Scutum with sparse punctuation, at least between notauli, distance between punctuations larger than the diameter of them. Scutellar foveae with very weak to effaced longitudinal carinae ( Fig. 8b ).......................................... 8 - Scutum with abundant punctuation, distance between punctuations similar to the diameter of them. Scutellar foveae with sharp longitudinal carinae ( Figs 10b & 11b )...................................................................... 9 8. Scutellar spine triangular. Scutellar foveae shallow, with effaced longitudinal carinae, nearly smooth ( Fig. 8b ). Occiput not clearly carinate.................................................................. A. jantonii Ros-Farré n. sp. - Scutellar spine elongated. Scutellar foveae deep, very slightly striate longitudinally ( Fig. 9b ). Occiput with thin but clear longitudinal carinae on dorsal half and with transverse carinae on ventral half..................... A. adelae Ros-Farré n. sp. 9. Scutellar foveae small and entirely carinate longitudinally ( Fig. 10b ). Scutellar spine narrow ( Fig. 10b )................................................................................. A. carlestolrai Ros-Farré & Pujade-Villar n. sp. - Scutellar foveae large, longitudinally carinate centrally (near interfoveal carina), smooth laterally ( Fig. 11b ). Scutellar spine triangular (wide anteriorly) ( Fig. 11b )................................... A. gemmae Ros-Farré & Pujade-Villar n. sp. 10. Scutellum very sharply convex, gibbous. Scutellar spine robust and blunt. End of scutellar disc uniformly narrowing to end of scutellar spine, so it is difficult to distinguish the beginning of scutellar spine. Scutellar foveae distinctly wide, scutellar pits sharply deep. Median mesoscutal furrow smooth and shining. Lateral line complete, reaching anterior margin of scutum ( Fig. 12a & b )......................................................................... A. enormis Belizin, 1952 - Scutellum flat, to boardly convex. Scutellar spine always pointed. End of scutellar disc different, the beginning of scutellar spine can always be distinguished. Character combination different............................................. 11 11. Scutellum convex, but sharply depressed on central area. Scutellar spine very short, 0.19 length of scutellar disc. Scutellar foveae wide, coriaceous anteriorly and slightly carinate posteriorly ( Fig. 13b )............... A. minutispina Belizin, 1952 - Scutellar disc never depressed on central area. Scutellar spine longer. Scutellar foveae variable, but never conspicuously coriaceous anteriorly...................................................................................... 12 12. Scutellar disc without interfoveal carina, weakly rugose. Scutellar spine long, 1.25 length of scutellar disc, and robust ( Fig. 14b ). Lateral face of pronotum with sharp transverse and sinuous carinae. Notauli conspicuously coriaceous and with abundant transverse carinae. Lateral line complete, reaching anterior margin of scutum. Area between notauli sharply prominent in lateral view ( Fig. 14a ).............................................................. A. elisendae Ros-Farré n. sp. - Scutellar disc with interfoveal carina (in A. daurica it is dificult to see), with variable sculpture. Scutellar spine shorter. Lateral face of pronotum with or without carinae. Notauli coriaceous or with transverse carinae, only in A. readae and A. sibirica with both characters toguether. Area between notauli flat or prominent............................................... 13 13. Scutum and scutellum shiny, without microsculpture, sometimes with macrosculpture ( Figs 15b ). Scutellum always clearly convex. Scutellar pits very deep. Interfoveal carina effaced on scutellar disc.................. A. abbreviata Belizin, 1952 - Scutum always coriaceous, in A. magdae very weakly so ( Fig. 39b ). Scutellum variable, flat to convex. Scutellar pits present or absent. Interfoveal carina present, except in A. daurica ....................................................... 14 14. Wing pubescence not reduced, marginal pubescence long, wing sharply smoked, with veins very conspicuously impressed (Rs+M present). Occiput with thin but noticeable carinae. Lateral surface of pronotum transversely carinate dorsally ( Fig. 16a ). Scutellar spine thin and slender. Scutellar disc with prominent interfoveal carina. Scutellar foveae with very weak to sharp transverse carinae anteriorly and longitudinal carinae posteriorly ( Fig. 16b )............... A. sergioi Ros-Farré n. sp. - Wing pubescence variable. Occiput variable, with or without carinae (but if it has noticeable carinae then lateral surface of pronotum is clearly rugose or with sharp transverse carinae). Lateral surface of pronotum coriaceous to carinate. Interfoveal carina variable. Scutellar foveae variable........................................................................ 15 15. Scutellar foveae with, at least, two sharp longitudinal carinae that continue, together with interfoveal carina, along scutellar disc. Scutellar pits sharply deep. Scutum with sharp coriaceous sculpture ( Figs 17b & 18b )......................... 16 - Scutellar foveae different, they can have weak longitudinal carinae except for A. daurica , which has sharp longitudinal carinae, but in this species interfoveal carina is effaced on scutellar disc ( Fig. 38b ). Scutellar pits present or absent. Scutum variable.................................................................................................... 17 16. Scutum with transverse carinae ( Fig. 17b )............................................. A. hartigi Dalla Torre, 1889 - Scutum coriaceous ( Fig. 18b )....................................... A. buffingtoni Ros-Farré & Pujade-Villar n. sp. 17. Scutellar disc with prominent interfoveal carina, (if there are lateral carinae, they are not so prominent). Carinae of occiput medium impressed ( Fig. 19b ). Lateral line complete ( Fig. 19a ). Marginal pubescence of wing long, exremely long in the dorsal margin between basis of wing and radial cell ( Fig. 3a )..................................... A. tomasi Ros-Farré n. sp. - Interfoveal carina variable. Marginal pubescence of wing different, never so long in dorsal margin. Character combination different.............................................................................................. 18 18. Occiput with sharp carinae ( Fig. 2c ). Lateral surface of pronotum coriaceous. Scutellum very slightly convex, nearly flat. Marginal pubescence of wing absent. Scutellar disc with a medium prominent interfoveal carina, which continues straight until half of length, and undulate posteriorly.............................................. A. utahensis Ashmead, 1896 - If occiput has sharp carinae, lateral surface of pronotum is clearly rugose or with sharp transversal carinae. Scutellum flat to convex. Marginal pubescence of wing variable. Scutellar disc with weak to prominent interfoveal carina............... 19 19. Occiput with or without weak carinae ( Figs 23b , 25b & 30b ). Lateral surface of pronotum coriaceous, rarely with scarce and weak transverse carinae on dorsal half ( Figs 23a , 25a & 30a ). Scutellar disc with a prominent interfoveal carina; lateral carinae of scutellar disc, if present, not so prominent as interfoveal carina ( Figs 23a , 25a , 31a & 31b )........................ 20 - Occiput with sharp to very sharp carinae ( Figs 39b & 40b ). Lateral surface of pronotum clearly rugose or with sharp transverse carinae ( Figs 40a , 42a & 48a ). Scutellar disc only with a noticeable, but not prominent, interfoveal carina; lateral carinae of scutellar disc are similar or weaker than interfoveal carina ( Figs 40a , 42a & b ).................................... 34 20. Lateral line complete, reaching anterior margin of scutum ( Figs 24a & b ). Marginal pubescence of wing long............ 21 - Lateral line incomplete, ending before anterior margin of scutum ( Figs 25a & b ). Marginal pubescence of wing variable... 24 21. Lateral surface of pronotum without transversal carinae ( Fig. 21a ). Scutellar foveae with weak longitudinal carinae posteriorly ( Fig. 21b ). Mesopleuron coarsely coriaceous on anterior 1/3................................. A. sibirica Kieffer, 1901 - Lateral surface of pronotum with a few weak transverse striae or carinae. Scutellar foveae smooth. Mesopleuron with clear punctate sculpture or with some rugose carinae on anterior 1/3................................................. 22 22. Scutellar spine short, 0.5 length of scutellar disc. Scutellar foveae sharply wide and shallow ( Fig. 22c ). Notauli transversely carinate and weakly alutaceous ( Fig. 22b ). Mesopleuron with clear punctate sculpture on anterior 1/3.................................................................................................... A. attenuata Belizin, 1952 - Scutellar spine long, 0.7 to 0.9 length of scutellar disc ( Figs 23b & 24b ). Scutellar foveae variable. Notauli coriaceous or alutaceous. ( Figs 23b & 24b ). Mesopleuron with some rugose carinae on anterior 1/3................................... 23 23. Notauli coriaceous. Median mesoscutal furrow wide anteriorly. Occiput coriaceous, with carinae ( Fig. 24b ). Area between notauli prominent in lateral view ( Fig. 24a ), pointed centrally. Scutellar disc rectangular, seen in lateral view at the same level than scutellar spine, which is flat........................................ A. martae Ros-Farré & Pujade-Villar n. sp. - Notauli smooth centrally, alutaceous near margins. Median mesoscutal furrow narrow dorsally. Occiput rugose behind ocelli, withouth carinae ( Fig. 23b ). Area between notauli not prominent in lateral view, curved. Scutellar disc oval, slightly sloping posteriorly towards basis of scutellar spine, which is directed downwards ( Fig. 23a )..................................................................................................... A. annae Ros-Farré & Pujade-Villar n. sp. 24. Scutellar disc very flat and wide, smoothly coriaceous and weakly rugose; lateral bar narrow and complete, scutelar disc entirely and sharply Marginated. Scutellar foveae very large. Scutum coriaceous. Median ridge weak near anterior margin of median mesoscutal furrow ( Fig. 25b ). Mesopleuron sharply coriaceous posterodorsally............................................................................................... A. marginata Ros-Farré & Pujade-Villar n. sp. - Scutellar disc and lateral barr different; sometimes marginate, rarely sharply or entirely, lateral bar never so narrow. Scutellar foveae usually smaller. Scutum variable. Median ridge different. Mesopleuron variable............................ 25 25. Scutellar foveae with one or two thin oblique carinae on each side of interfoveal carina. Scutellar spine extremely short. Notauli and median mesoscutal furrow transversely striate ( Fig. 26b )....... A. santamariai Ros-Farré & Pujade-Villar n. sp. - Scutellar foveae different. Character combination different.................................................... 26 26. Scutellar foveae rounded, transversely carinate anteriorly and rugose posteriorly, deep and small, lateral bar (between scutellar fovea and scutellum margin) very wide. Scutellar spine short, equilateral-triangle-shaped. ( Fig. 27b ). Ocelli clearly prominent................................................................... A. mireiae Ros-Farré & Pujade-Villar n. sp. - Scutellar foveae different, lateral bar not as wide as in A. mireiae . Scutellar spine usually longer than wide ( Figs 28b & 29b ). Ocelli slightly prominent............................................................................... 27 27. Scutellar disc not swollen posterior to scutellar foveae ( Figs 28a , 29a & 31a )...................................... 28 - Scutellum swollen posterior to scutellar foveae (35a), sometimes very slightly so, in this case marginal pubescence of wing very noticeable...................................................................................... 31 28. Lateral frontal carinae absent on dorsal half, near ocelli, in this area there is a clear and abundant punctate sculpture ( Fig. 1e )............................................................... A. punctifrons Ros-Farré & Pujade-Villar n. sp. - Lateral frontal carinae complete, punctuations never so sharp ( Figs 1f & 1d )...................................... 29 29. Scutellar spine short and wide. Scutum slightly punctate and coriaceous ( Fig. 29b )............. A. teresae Ros-Farré n. sp. - Scutellar spine long and thin. Scutum coriaceous, with abundant to scarce transverse carinae. ( Figs 30b & 31b ).......... 30 30. Scutellum sloping downwards on each side of interfoveal carina (roof-shaped), which is extremely prominent and sharp. Scutellar disc sharply rugose and with some sinuous longitudinal carinae ( Fig. 30b )................................................................................................... A. carinata Ros-Farré & Pujade-Villar n. sp. - Scutellum flat on each side of interfoveal carina, which is prominent but fine. Scutellar disc rugose, without longitudinal carinae after foveae ( Fig. 31b )........................................................... A. dianae Ros-Farré n. sp. 31. Marginal pubescence of wing present, uniformly distributed in all the wing margin................................. 32 - Marginal pubescence of wing absent or discontinuous........................................................ 33 32. Notauli narrow. Scutellum with sharp longitudinal carinae. Scutellar foveae coriaceous on posterior 1/3 ( Fig. 32b ).............................................................................. A. julii Ros-Farré & Pujade-Villar n. sp. - Notauli wide. Scutellum with some fine longitudinal carinae. Scutellar foveae with weak carinae ( Figs 33a & b )............................................................................................ A. proxima Belizin, 1952 33. Scutum very coriaceous. Scutellar foveae shallow, with abundant transverse carinae on anterior half and scarce longitudinal carinae on posterior half ( Fig. 34b )...................................... A. kiefferi Ros-Farré & Pujade-Villar n. sp. - Scutum with short transverse carinae, mostly between notauli. Scutellar foveae deep and rugose ( Fig. 35b )................................................................................ A. zuparcoi Ros-Farré & Pujade-Villar n. sp. 34. Scutellar disc convex after foveae ( Figs 37a , 38b & 39 ) Marginal pubescence of wing variable....................... 35 - Scutellar disc flat after foveae ( Figs 44a & 45b ). Marginal pubescence of wing starting posterior to R2, conspicuous...... 42 35. Lateral line complete, reaching anterior margin of scutum. Wing pubescence not reduced, marginal pubescence long and dense. Scutum without transverse carinae.................................................................. 36 - Character combination different.......................................................................... 37 36. Scutellar foveae weakly sculptured and shallow ( Fig. 36b ). Scutellar disc sharply convex.......... A. porif Ros-Farré n. sp. - Scutellar foveae smooth and deep ( Fig. 37b ). Scutellar disc weakly convex................. A. simulacrum Belizin, 1952 37. Punctuations of 4 th metasomal segment reaching posterior margin of 3 rd segment in dorsal view. Interfoveal carina not continuing on scutellar disc. Scutum with scarce microsculpture ( Fig. 38b ). Occiput with transverse carinae, even behind ocelli. Wing membrane hyaline.................................................................. A. daurica Belizin, 1952 - Punctuations of 4 th metasomal segment not reaching posterior margin of 3 rd segment in dorsal view. Interfoveal carina present on scutellar disc ( Fig. 39b ). Character combination different................................................... 38 38. Lateral line incomplete, only reaching basal 2/3 of scutum ( Fig. 39b & 40a )...................................... 39 - Lateral line complete, reaching anterior margin of scutum ( Figs 43a & b ); in A. sculpturata , although they are very conspicuous they end just before anterior margin of scutum ( Fig. 42a ).................................................. 40 39. Scutellar foveae with weak oblique carinae, scutellar pits deep. Scutum very slightly coriaceous. Occiput with short longitudinal carinae behind ocelli ( Fig. 39b ). Lateral surface of pronotum entirely carinate transversely ( Fig. 39a )................................................................................... A. magdae Ros-Farré & Pujade-Villar n. sp - Scutellar foveae smooth, scutellar pits absent. Scutum with noticeable coriaceous sculpture. Occiput with long longitudinal carinae behind ocelli ( Fig. 40b ). Lateral surface of pronotum transversely carinate dorsally and ventrally, coriaceous centrally ( Fig. 40a ).......................................................... A. belizini Ros-Farré & Pujade-Villar n. sp. 40. Scutum with noticeable rugose carinae ( Fig. 42b ). Notauli sharply carinate transversely............................ 41 - Scutum usually coriaceous, but sometimes with weak and short transverse carinae ( Fig. 41b ). Notauli weakly carinate transversely................................................................ A. suecica Dalla Torre & Kieffer, 1910 41. Scutellar pits deep, scutellum rugose ( Fig. 42b ). Lateral line nearly reaching anterior margin of scutum ( Fig. 42a ). ......................................................................................... A. scutellata ( Villers, 1789 ) - Scutellar pits absent, scutellum with parallel longitudinal carinae ( Fig. 43b ). Lateral line complete, reaching anterior margin of scutum ( Fig. 43a )................................................................. A. blancae Ros-Farré n. sp. 42. Subpronotal plate conspicuous and projected ahead ( Fig. 44b ). Frontal and lateral frontal carina very conspicuous ( Fig. 1a ). Scutellar spine extremely robust, wide and long, scutellar foveae noticeably rugose, scutellar pits sharply deep ( Fig. 44b )................................................................... A. forshzarai Pujade-Villar & Ros-Farré n. sp. - Subpronotal plate not projected or only projected basally ( Fig. 45b ). Lateral frontal carinae variable, frontal carinae absent or medium impressed ( Fig. 1c ). Scutellar spine never so robust, wide and long, scutellar foveae smooth, with weak longitudinal carinae or slightly alutaceous, scutellar pits absent ( Figs 45b , 46b & 48b )........................................ 43 43. Lateral frontal carinae, parallel, straight with facial carinae, which are also parallel ( Fig. 1c ). Notauli coriaceous with scarce transverse carinae ( Fig. 45b )........................................... A. readae Ros-Farré & Pujade-Villar n. sp. - Lateral frontal carinae, confluent towards antennal foramina, not straight with facial carinae, which are divergent ( Fig. 1b ). Notauli shiny, with transverse carinae..................................................................... 44 44. Scutellar spine robust, wide basally, medium length (0.55 to 0.65 length of scutellar disc). Scutellar disc with sharp, prominent and sinuous longitudinal carinae ( Fig. 46b )............................................ A. caminali Ros-Farré n. sp. - Scutellar spine slender, narrow basally, long (0.65 to 1.20 length of scutellar disc). Scutellar disc different ( Fig. 47b , 48b & 49b )................................................................................................ 45 45. Scutellar foveae shallow, rugose, scutellar disc rugose ( Fig. 47b ).............. A. robusta Ros-Farré & Pujade-Villar n. sp. - Scutellar foveae deeper, smooth or with effaced sculpture, scutellar disc with straight longitudinal carinae ( Figs 48b & 49b ). (species difficult to separate)............................................................................ 46 46. Robust insect (males>3.0mm., females> 3.4mm .). Scutum with or without weak transverse carinae, notauli and median mesoscutal furrow with small and weak transverse carinae ( Fig. 48b ). Interfoveal carina continuing along scutellar disc to beginning of scutellar spine................................................................. A. longispina Kieffer, 1901 - Insect not robust (males< 2.7mm ., females unknown). Scutum with sharp transverse carinae, notauli and median mesoscutal furrow with sharp transverse carinae ( Fig. 49b ). Interfoveal carina continuing along scutellar disc and spine until last 1/3 of scutellar spine................................................... A. danielssoni Ros-Farré & Pujade-Villar n. sp.