3531 Author Perkins, Philip D. text Zootaxa 2012 2012-10-30 3531 1 95 journal article 1175­5334 C1FA90AF-1C31-45D6-9CB6-C7D3058E501C Epimetopus lacordairei Orchymont, 1933 Figs. 10 (habitus), 14 (aedeagus), 127 (map) Epimetopus lacordairei Orchymont, 1933: 288 . Type Material. Holotype (male, greatly disarticulated): Brazil : "Matto Grosso, Corumba" ( ISNB ). New collection records: Bolivia : Beni , Ilha Flores , Rio Itenez , 13° 2' S , 62° 43' W , 7 viii 1964 , J. Bouseman & J. Lussenhop (5 MCZ ) ; Paraguay : Dep. Concepcion , Arroyo Toro Paso , 23° 51' S , 56° 46' W , 25 x 2002 , U. Drechsel (5 NMW ) ; Arroyo Toro Paso , Unterlauf (contrib. Rio Apa ), seepage, 23° 51' S , 56° 46' W , 24 x 2002 , U. Drechsel (102 NMW ) ; Stillwasser des Arroyo Toro Paso , Seitenam ( Uferschlamm ), 23° 51' S , 56° 46' W , 25 x 2002 , U. Drechsel (8 NMW ) . Differential diagnosis. Differentiated from all members of the Costatus group, except E. trilobus , by the trilobed apical margin of the pronotal hood ( Figs. 10 , 32 ). The aedeagus of E. lacordairei ( Fig. 14 ), with an apically deeply bifid median lobe, differs markedly from all other members of the genus. The aedeagus of E. trilobus ( Fig. 36 ) probably represents a preliminary stage in the evolution of the extreme condition seen in E. lacordairei . This view is supported by the fact that both species have a tri-lobed pronotal apex—the only species now known to have that condition. Distribution. Currently known from Bolivia , Brazil , and Paraguay ( Fig. 127 ).