Revision of theBaeoglena species of the West Palaearctic Region (Coleoptera: Staphylinidae: Aleocharinae) Author Assing, Volker text Beiträge Zur Entomologie = Contributions to Entomology 2019 2019-06-24 69 1 1 32 journal article 10.21248/contrib.entomol.69.1.001-032 0005-805X 5742658 016AAB3F-8AE2-4759-B95A-DEAA074768CD Oxypoda ( Baeoglena ) caucasica BERNHAUER, 1902 ( Figs 55–58 , Maps 1 , 7 ) Oxypoda ( Baeoglena ) nova BERNHAUER, 1902: 180 . Type material examined : Lectotype , present designation: “Caucasus, Helenendorf, Reitter. / fusina Rey / vid. Bernh. / caucasica Bh. / ex coll. Skalitzky / Lectotypus Oxypoda caucasica Bernhauer , desig. V. Assing 2018 / Oxypoda caucasica Bernhauer , det. V. Assing 2018” ( NHMW ). Comment : The original description is based on an unspecified number of syntypes from “Caucasus: Helenendorf, Martkopi, Meskisches Gebige [sic], Daghestan, Lenkoran ” ( BERNHAUER 1902 ). A male syntype from “Helenendorf ” is designated as the lectotype . Material examined : Ukraine : 1, 1 ex. , Odessa obl. , Dobroolexandrivka [“Alexanderhilf ”; 46°20'N , 30°29'E ], leg. Leder ( NHMW ) . Turkey : Bolu : 1, 1, road Bolu – Yığılca , 40°51'N , 31°39'N , 1060 m , 29.IV.2012 , leg. Brachat & Meybohm (cAss); 5, 5, 23 km S Bolu , 40°38'N , 31°37'N , 1540 m , 30.IV.2012 , leg. Brachat & Meybohm (cAss); 1, Karabük , pass W Eskipazar , 40°58'N , 33°23'E , 1400 m , beech forest, litter sifted, 26.III.2010 , leg. Assing (cAss) . Kastamonu : 1, 5 exs. [found together with O. nova ], 5 km S Küre , 1200 m , 18.V.1976 , leg. Besuchet & Löbl ( MHNG , cAss) . Sinop : 1, 2, Çangal Dağı , 7–15. VI.1960 , leg. Schubert (cAss) . Trabzon : 1, ca. 50 km S Trabzon , 20 km S Maçka , Altındere Milli Park , 40°40'N , 39°40'E , 1560 m , spruce forest with Rhododendron , sifted, 26.VII.2006 , leg. Schülke ( MNB ) . Rize : 1, ca. 50 km S Ardeşen , Çat , 40°52'N , 40°56'E , 1240 m , Alnus forest, sifted, 3.VIII.2006 , leg. Schülke ( MNB ); 1, ca. 30 km SW Hopa , river valley of Çağlayan D., ca. 41°15'N , 41°13'E , 500 m , 29.VI.1998 , leg. Solodovnikov (cAss) . Artvin : 6 exs. , Borçka , 1–3.VI.1960 , leg. Schubert (cAss); 5 exs. , pass between Borçka and Hopa , 700 m , 8.VI.1986 , leg. Besuchet , Löbl & Burckhardt ( MHNG , cAss); 1 ex. , same locality, 14.V.1967 , leg. Besuchet ( MHNG ); 68 exs. , Karkal Dağı , Pirnalli , 1600 m , 11.VI.1986 , leg. Besuchet , Löbl & Burckhardt ( MHNG , cAss); 14 exs. , Karkal Dağı , below Pirnalli , 1250 m , 11.VI.1986 , leg. Besuchet , Löbl & Burckhardt ( MHNG , cAss); 3 exs. , S Artvin , 800 m , 7.VI.1986 , leg. Besuchet , Löbl & Burckhardt ( MHNG ); 19 exs. , S Artvin , 1500 m , 9.VI.1986 , leg. Besuchet , Löbl & Burckhardt ( MHNG , cAss); 41 exs. , SW Artvin , 1900 m , 9.VI.1986 , leg. Besuchet , Löbl & Burckhardt ( MHNG , cAss) . Kars : 1, Kars , 1750 m , 17.VI.1986 , leg. Besuchet , Löbl & Burckhardt ( MHNG ) . Russia : 1 ex. , Krasnodar , 35 km NNE Sochi, Babuk-Aul, 43°53'N , 39°49'E , 560 m , Fagus orientalis and Castanea sativa forest, litter and bark sifted, 11.VII.2011 , leg. Assing (cAss); 2 exs. , Krasnodar , Temnolesskaia, near Mezmay, 800 m , 8.VI.1999 , leg. Smetana (cAss); 25 exs. [partly teneral], Karachayevo-Cherkesskaya Respublika, 13 km SW Teberda, 43°20'N , 41°40'E , 1450 m , moist spruce forest with scattered beech, litter, moss, and dead wood sifted, 22.VII.2011 , leg. Assing (cAss); 10 exs. [partly teneral], Karachayevo-Cherkesskaya Respublika, 20 km SW Teberda, above Dombai, 43°17'N , 41°38'E , 1950 m , mixed forest (fir, spruce, beech), leaf litter sifted, 25.VII.2011 , leg. Assing (cAss); 1 ex. , Karachayevo-Cherkesskaya Respublika, 9 km SW Teberda, Teberdinski range, Baduk river valley, 43°23'N , 41°40'E , 2000 m , subalpine forest (maple, birch, with rhododendron) near lakeshore, litter sifted, 26.VII.2011 , leg. Assing (cAss); 11 exs. , Karachayevo-Cherkesskaya Respublika, 9 km SW Teberda, Teberdinski range, Baduk river valley, 43°23'N , 41°40'E , 2000 m , spruce forest, bark of spruce and maple sifted, 27.VII.2011 , leg. Assing (cAss); 1 ex. , Krasnaja Poljana, Mt. Atchichkho, 800 m , 26.VI.1989 , leg. Arndt ( SDEI ); 2 exs. , Krasnaja Poljana, leg. Roubal ( NHMW ). Georgia : 6 exs. , Algeti National Park , W Manglisi , 41°42'N , 44°21'E , 1120 m , 11.VII.2015 , leg. Brachat & Meybohm (cAss) ; 6 exs. , Algeti National Park , W Manglisi , 41°42'N , 44°19'E , 1110 m , 11.VII.2015 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Algeti National Park , W Manglisi , 41°42'N , 44°18'E , 1210 m , 11.VII.2015 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Algeti National Park , Manglisi , 41°42'N , 44°23'E , 1190 m , 12.VII.2015 , leg. Brachat & Meybohm (cAss) ; 10 exs. , same data, but 13.VII.2015 (cAss) ; 1 ex. , Central Caucasus , Schatili –Mutso, 42°40'N , 45°11'E , 1390 m , 15.VII.2015 , leg. Brachat & Meybohm (cAss) ; 5 exs. , Central Caucasus , Gudani , 42°32'N , 44°59'E , 1710 m , 18.VII.2015 , leg. Brachat & Meybohm (cAss) ; 4 exs. , Gudani–Zhinvali , 42°27'N , 44°56'E , 1200 m , 19.VII.2015 , leg. Brachat & Meybohm (cAss) ; 8 exs. , Gudani–Zhinvali , 42°27'N , 44°56'E , 1200 m , 19.VII.2015 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Shida Kartli , Kvishkheti , 41°58'N , 43°30'E , 810 m , 12.V.2016 , leg. Brachat & Meybohm (cAss) ; 6 exs. , Shida Kartli , Kvishkheti , 41°57'N , 43°30'E , 790 m , 12.V.2016 , leg. Brachat & Meybohm (cAss) ; 6 exs. , Samtskhe-Javakheti , Timotesubani , 41°49'N , 43°31'E , 1144 m , 13.V.2016 , leg. Brachat & Meybohm (cAss) ; 3 exs. , Samtskhe-Javakheti , Bakuriani , 41°44'N , 43°43'E , 1770 m , 13.V.2016 , leg. Brachat & Meybohm (cAss) ; 2 exs. , Samtskhe-Javakheti , N Abastumani , 41°46'N , 42°50'E , 1370 m , 15.V.2016 , leg. Brachat & Meybohm (cAss) ; 2 exs. , Samtskhe-Javakheti , W Abastumani , 41°45'N , 42°49'E , 1345 m , 15.V.2016 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Abastumani , leg. Leder ( NHMW ) ; 3 exs. , Racha , 4 km N Nakerala , 42°24'N , 43°02'E , 1150 m , 18.V.2016 , leg. Brachat & Meybohm (cAss) ; 8 exs. , Shida Kartii , Kvishkheti , 41°57'N , 43°29'E , 1300 m , 24.VII.2016 , leg. Meybohm (cAss) ; 1 ex. , Zemo Svaneti , 4 km N Mazeri , 43°06'N , 42°36'E , 1690 m , 28.VII.2016 , leg. Meybohm (cAss) ; 2 exs. , Zemo Svaneti , 2 km N Ipari , 43°01'N , 42°50'E , 1670 m , 29.VII.2016 , leg. Meybohm (cAss) ; 1 ex. , Adjara , Skhalta valley , 41°35'N , 42°22'E , 800 m , 22.VI.2017 , leg. Brachat & Meybohm (cAss) ; 4 exs. , Adjara , Skhalta valley , 41°34'N , 42°25'E , 970 m , 22.VI.2017 , leg. Brachat & Meybohm (cAss) ; 3 exs. , Svaneti , Mazeri , 43°06'N , 42°36'E , 1660 m , 26.VI.2017 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Svaneti , Mestia–Hatsvali , 43°01'N , 42°45'E , 2350 m , 27.VI.2017 , leg. Brachat & Meybohm (cAss) ; 4 exs. , Svaneti , Mestia–Ushguli , 3 km N Ipari , 43°01'N , 42°50'E , 1750 m , 28.VI.2017 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Svaneti , Mestia–Ushguli , Kala , 42°57'N , 42°53'E , 1680 m , 28.VI.2017 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Imereti , Baghdati , 42°01'N , 42°49'E , 290 m , 18.V.2018 , leg. Brachat & Meybohm (cAss) ; 3 exs. , Imereti , S Sairme , 41°51'N , 42°47'E , 1930 m , 19.V.2018 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Imereti , S Sairme , 41°52'N , 42°47'E , 1890 m , 19.V.2018 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Imereti , S Sairme , 41°53'N , 42°46'E , 1420 m , 20.V.2018 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Imereti , S Sairme , 41°53'N , 42°46'E , 1360 m , 20.V.2018 , leg. Brachat & Meybohm (cAss) ; 6 exs. , Racha , E Shovi , 42°41'N , 43°42'E , 1720 m , 28.V.2018 , leg. Brachat & Meybohm (cAss) ; 3 exs. , Racha , E Shovi , 42°41'N , 43°42'E , 1700 m , 28.V.2018 , leg. Brachat & Meybohm (cAss) ; 2 exs. , Racha , W Glola , 42°42'N , 43°36'E , 1170 m , 28.V.2018 , leg. Brachat & Meybohm (cAss) ; 1 ex. , Ananuri Forest , 8.V.2006 , leg. Chalandze (cAss) ; 1 ex. , Trialetskiy Khrebet , Bakuriani , 1500–1600 m , 8.VII.1986 , leg. Wrase & Schülke ( SDEI ) ; 1 ex. , Tbilisi env., Zchneti , 800 m , VI.1987 , leg. Wrase & Schülke ( SDEI ) ; 2 exs. , 60 km N Tbilisi , Dusheti , 1500 m , 25.IX.1987 , leg. Oehlke ( SDEI ) ; 3 exs. , Khashuri [ 41°59'N , 43°35'E ] ( NHMW ) ; 1 ex. , “ Swanetien ”, leg. Leder ( NHMW ) ; 6 exs. , Likhskiy Khrebet [“Meskisches Geb.”], leg. Leder ( NHMW ) . Armenia : see ASSING & SCHÜLKE (2019) . Nagorno-Karabakh : see ASSING & SCHÜLKE (2019) . Azerbaijan : 2 exs. , Lenkoran , leg. Leder ( NHMW ) ; 1 ex. , Lerik , leg. Leder ( NHMW ) ; 4 exs. , Talysh range ( MHNG , NHMW ) . Iran : Mazandaran : 1 ex. , Elburz Mts. , Ramsar Co. , Eshkatechal , 36°51'N , 50°33'E , 1050 m , small stream, sifted, 6.VI.2008 , leg. Pütz (cAss) ; 1 ex. , Elburz Mts. , Babol Co. , 2 km SW Firuz Jah , 36°11'N , 52°39'E , 840 m , small stream, 31.V.2008 , leg. Pütz (cAss) ; 2 exs. , Elburz Mts. , Tonekabon Co. , 4.5 km SW Khanian , Sehezar forest , 36°33'N , 50°50'E , 940 m , small stream, litter and debris sifted, 5.VI.2008 , leg. Pütz (cAss) ; 4 exs. , Elburz Mts. , Sari Co. , Mohammabad , NE Sangdeh , 36°04'N , 53°10'E , 1530 m , beech forest, litter sifted, 29.V.2008 , leg. Pütz (cAss) . 1 ex. , Elburz Mts. , Sari Co. , Mohammabad , E Qolqol , 36°10'N , 53°10'E , 920 m , litter sifted, 29.V.2008 , leg. Pütz (cAss) ; 1 ex. , Elburz Mts. , Chalus Co. , SE Abbasabad , 36°39'N , 51°10'E , 280 m , leaf litter and debris sifted, 4.VI.2008 , leg. Pütz ( MNB ) ; 3 exs. , Taquiabad , 24.VII.1977 , leg. Huber ( MHNG ) . Gilan : 1 ex. , Elburz Mts., Rasht Co., 29 km S Rasht, 37°01'N , 49°37'E , 140 m , sifted, 7.VI.2008 , leg. Pütz (cAss). Locality ambiguous, not identified, or not specified : 1 ex. , “Armen. Geb.”, leg. Leder ( NHMW ) ; 6 exs. , “Kaukas”, “Caucasus”, etc., leg. Leder , Reitter , etc. ( NHMW ) ; 1 ex. , “Kaukas, ... Urwald bei Tcharsach”, 1.V.1875 , leg. Leder ( MHNG ) . Diagnosis : Winged species of mostly dark and more rarely reddish coloration. : median lobe of aedeagus ( Figs 55–56 ) approximately 0.35 mm long; ventral process weakly curved in lateral view, apically truncate in lateral view and convex in ventral view; crista apicalis long and broad; internal sac with pair of fork-shaped apical structures (lateral view) and with a weakly curved and weakly sclerotized flagellum; apical portion of paramere ( Fig. 57 ) relatively short, barely half as long as basal portion, rather weakly dilated basally. : spermatheca ( Fig. 58 ) with minute apical cuticular invagination. Distribution and natural history : The distribution of O. caucasica ranges from South Ukraine and Northwest Anatolia across North Anatolia , the Greater Caucasus, and the Lesser Caucasus (including adjacent regions) to North Iran ( Maps 1 , 7 ). The record from Germany in SCHÜLKE & SMETANA (2015) is based on LOHSE (1989: 231) “nach Angaben von L. Zerche”, who reports a male from the environs of Gotha collected in 1867. This specimen was clearly either mislabelled or misidentified, as can be inferred from the revised distribution of O. caucasica . The examined material was sifted from leaf litter, roots, moss, and bark in various forest (deciduous, coniferous, mixed) habitats and near streams. The altitudes range from 140 to 2350 m . The specimens were found in March (one record), April (2), May (26), June (27), July (37), August (1), and September (1).