Leiocassis rudicula (Teleostei: Siluriformes: Bagridae), a new catfish from eastern Borneo Author Ng, Heok Hee Author Hadiaty, Renny K. text Zootaxa 2019 2019-08-02 4651 1 191 200 journal article 26114 10.11646/zootaxa.4651.1.12 a816c380-3db4-4a65-a74e-83a794dd51ee 1175-5326 3358940 E38FEDA7-AA93-4A5C-8AEF-A50A8E1A7860 Key to the species of Leiocassis (modified from Ng & Tan, 2018 ) 1. Posterior tip of supraoccipital process in contact with nuchal plate; anal fin with broad brown band covering at least middle third................................................................................................ 2 - Posterior tip of supraoccipital process not in contact with nuchal plate; anal fin with irregular narrow brown band covering less than middle third...................................................................................... 3 2. Length of adipose-fin base 16–17% SL; dorsal-spine length 21% SL; pectoral-spine length 18–19% SL [ Sumatra : Alas River drainage].................................................................................... L. aculeata - Length of adipose-fin base 13–15% SL; dorsal-spine length 16–18% SL; pectoral-spine length 15–18% SL [Malay Peninsula: Mae Nam Kolok drainage southwards to Endau River drainage; Sumatra : Wampu River drainage southwards to Musi River drainage; Java : Ciberang River drainage; Borneo: Kapuas River drainage]............................. L. poeciloptera 3. Body uniform medium brown, without cream or yellow irregular bands or patches; pectoral-spine length 9–13% SL [Borneo: Kayan River drainage]........................................................................ L. tenebrica - Body mottled brown, with cream or yellow irregular bands or horizontally elongate patches; pectoral-spine length 13–21% SL................................................................................................. 4 4. Posterior margin of dorsal spine smooth................................................................... 5 - Posterior margin of dorsal spine with 5–22 serrae............................................................ 6 5. Horizontally elongate cream or yellow patches on sides of body between dorsal and adipose fins; outermost principal caudal-fin rays with short extensions; body depth at anus 11–16% SL; caudal peduncle depth 6–7% SL [Borneo: Mentaya, Sebangau and Kahayan river drainages; Sumatra : Lalang and Batang Hari river drainages]............................... L. bekantan - Irregular cream or yellow band on sides of body between dorsal and adipose fins; outermost principal caudal-fin rays without extensions; body depth at anus 16–20% SL; caudal peduncle depth 9–11% SL [Borneo: Mahakam River drainage].......................................................................................... L. rudicula new species 6. Length of adipose-fin base 19–22% SL; head length 26–29% SL [Borneo: Kalabakan and Segama river drainages]. L. collina - Length of adipose-fin base 14–18% SL; head length 29–34% SL [Bangka: Baturusa River drainage; Belitung: Cerucuk River drainage; Borneo: Kapuas River drainage eastwards and northwards to Padas River drainage, Barito River drainage]............................................................................................... L. micropogon