New taxa of pygmephoroid mites (Acari: Pygmephoroidea: Pygmephoridae, Neopygmephoridae) phoretic on Enoplotrupes sharpi (Coleoptera: Geotrupidae) from Thailand Author Khaustov, Alexander A. Author Frolov, Andrey V. text Zootaxa 2018 2018-07-02 4442 2 277 292 journal article 29764 10.11646/zootaxa.4442.2.5 0fea0081-d093-4611-89c2-316f9baf6ca4 1175-5326 1303125 8EB2105E-3EAA-444A-A20E-819AD215B3C3 Key to world species of Spatulaphorus (based on Khaustov & Trach 2018 ) 1. Genu I with two setae.................................................................................. 2 - Genu I with one seta................................................. S. copridis Khaustov, 2007a ( Crimea , Iran ) 2. Tarsi II and III each with spiniform setae pl” and tc’ ......................................................... 3 - Tarsi II and III without spiniform setae, or only pl” or tc’ spiniform.............................................. 5 3. Setae sc 2 foliate.............................................. S. vladimiri Khaustov, 2005 ( Ukraine , Crimea , Iran ) - Setae sc 2 not modified, setiform.......................................................................... 4 4. Setae c 2 foliate, blunt-ended...................................... S . venustus Khaustov and Trach, 2012a ( Ukraine ) - Setae c 2 not modified, pointed................................... S. altaicus Khaustov and Trach, 2018 ( Russia : Altai ) 5. At least setae ( pv ) on tarsus II foliate distally................................................................ 6 - Setae ( pv ) on tarsus II not foliate distally.................................................................. 10 6. Tibiotarsus I with large, hook-like claw and a distinct counterpart opposite to the claw base........................... 7 - Tibiotarsus I with a small, almost straight claw, and no counterpart......... S. foliatus Dastych and Rack, 1993 ( Botswana ) 7. Tarsi II and III with setae tc’ not modified.................................................................. 8 - Tarsi II and III with setae tc’ spiniform................................... S. langi Dastych and Rack, 1993 ( Vietnam ) 8. Setae ps 3 subequal to or shorter than ps 1, 2.................................................................. 9 - Setae ps 3 distinctly longer than ps 1, 2.......................... S. luriei Dastych, Rack & Camerik, 1997 ( South Africa ) 9. Setae ( pv ) on tarsus III foliate, setae ps 1, 2 distinctly barbed....................... S. enoplotruporum sp. nov. ( Thailand ) - Setae ( pv) on tarsus III not foliate, setae ps 1, 2 smooth.................. S. camerikae Dastych and Rack, 1993 ( Botswana ) 10. Tarsi II and III with setae pl” not modified................................................................. 11 - Tarsi II and III with setae pl” spiniform.................. S. gorganica Rahiminejad and Hajiqanbar, 2011 ( Iran , Crimea ) 11. Apodemes 4 long, exceeding beyond setae 3 b .............................................................. 12 - Apodemes 4 vestigial................................................. S. geotruporum Khaustov, 2005 ( Ukraine ) 12. Setae e and h 2 subequal................................................................................ 13 - Setae e more than two times longer than h 2.................................. S. geotrupi ( Mahunka, 1970 ) ( Hungary ) 13. Setae e and h 2 barbed; setae ps 3 more than two times longer than ps 1, 2..... S. gymnopleuri Sobhi and Hajiqanbar, 2017 ( Iran ) - Setae e and h 2 smooth; setae ps 3 subequal to ps 1, 2..................... S . braziliensis Khaustov and Frolov, 2017 ( Brazil )