A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini) Author Caterino, Michael S. Author Tishechkin, Alexey K. text ZooKeys 2013 271 1 401 http://dx.doi.org/10.3897/zookeys.271.4062 journal article http://dx.doi.org/10.3897/zookeys.271.4062 1313-2970-271-1 Operclipygus lunulus sp. n. Figs 30G32 H-KMap 11 Type locality. PANAMA: Colon : San Lorenzo National Park, Achiote [ 9°12'N , 79°59'W ]. Type material. Holotype male: "PANAMA: Prov. Colon , Achiote - P. N. San Lorenzo N09°12', W79°59' Cafetal B Dist. 50m Tr. Intercepcion , A. Mercado 8 -22.V.2007" / " Operclipygus sp. # 12 San Lorenzo Inventory A.K. Tishechkin det. 2010" / "Caterino/Tishechkin Exosternini Voucher EXO-00282" (FMNH). Paratype (1): same data as type (GBFM). Diagnostic description. This species is extremely similar to Operclipygus dubius , differing mainly in the following characters: length: 1.59-1.62 mm, width: 1.22-1.28 mm; central portion of frontal stria present, detached from sides; supraorbital and frontal striae united in a broad, continuous arch; recurved arms of anterior submarginal stria ending near median pronotal gland openings, about 8 puncture widths from anterior margin; short fragment of lateral submarginal pronotal stria present in anterior corner; prescutellar impression slightly shorter than scutellum; prosternal striae divergent posteriorly, not connected at base; mesometaventral stria arching narrowly forward at middle; propygidium with moderate, round, shallow punctures separated by about their widths or slightly less basally, less dense posterad; pygidium with coarser punctures sparsely scattered in basal third of disk; pygidial sulcus complete, deep, strongly crenulate along outer edge. Male genitalia (Figs 32 H-K ) as for group description, with the apices of S8 bluntly subtruncate, dorsal edge oblique to apex, bearing about 5 long apical setae, with distinct submarginal sclerotization along upper margins; T9 with apices obliquely subquadrate, expanded to inner edges; T10 not divided; S9 desclerotized along midline, strongly expanded to base, shallowly but acutely emarginate at apex; tegmen more or less parallel-sided, rounded to apex, very similar to that of Operclipygus andinus , but distinctly more elongate. Remarks. The short recurved arms of the anterior submarginal stria (Fig. 30G) combined with the presence of a frontal stria and strongly reduced lateral submarginal pronotal striae distinguish this species from several close relatives. Etymology. The name of this species refers to the continuous, crescent-like arch formed by the frontal and supraorbital striae.