<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2022" docId="F895E5960025550FA394CC1243A1DE43" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina aureola Stoliczka 1872" docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-08-25</mods:number>
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<mods:number>1119</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<treatment ID-GBIF-Taxon="199839000" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43" httpUri="http://treatment.plazi.org/id/F895E5960025550FA394CC1243A1DE43" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="F895E596-0025-550F-A394-CC1243A1DE43" authority="Stoliczka, 1872" authorityName="Stoliczka" authorityYear="1872" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">Pupina aureola Stoliczka, 1872</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Umbilical, columellar and parietal views of A Pupinella mansuyi, specimen CUMZ 12148 from Pha Chu, Nan B Pupina artata from the Pupina artata species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai C Pupina godwinausteni sp. nov. from the Pupina arula species group, holotype CUMZ 12090 / 1 D Pupina aureola from the Pupina aureola species group, specimen CUMZ 12130 from Sra Morakot, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1">Figs 10D</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Opercula of three mainland Southeast Asian Pupina species groups A Pupina bensoni sp. nov. from the Pupina artata species group, specimen CUMZ 12047 B Pupina siamensis from the Pupina arula species group, specimen CUMZ 12067, and C Pupina aureola from the Pupina aureola species group, specimen CUMZ 12116. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure19" httpUri="https://binary.pensoft.net/fig/735380" pageId="0" pageNumber="1">, 19C</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">, 32G, H</figureCitation>
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 &quot; forma minor &quot; from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B)." figureDoi="10.3897/zookeys.1119.85400.figure34" httpUri="https://binary.pensoft.net/fig/735395" pageId="0" pageNumber="1">, 34F</figureCitation>
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">, 36A-F</figureCitation>
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G)." figureDoi="10.3897/zookeys.1119.85400.figure38" httpUri="https://binary.pensoft.net/fig/735399" pageId="0" pageNumber="1">, 38A-E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Stoliczka" authorityYear="1872" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">Pupina aureola</taxonomicName>
Stoliczka, 1872: 267, pl. 10, figs 11, 12. Type locality: Penang [Penang State, Malaysia]. 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Indian Museum, Calcutta" pageId="0" pageNumber="1" refId="B104" refString="Nevill, G, 1878. Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Indian Museum, Calcutta, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
: 299. de Morgan 1885: 414, Poulo Pinang, mont Tchora, 
<normalizedToken originalValue="près">pres</normalizedToken>
<normalizedToken originalValue="d’Ipoh">d'Ipoh</normalizedToken>
(Kinta), [Perak State, Malaysia]. 
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" author="von Moellendorff, OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="330 - 348" refId="B191" refString="von Moellendorff, OF, 1891. On the land and freshwater shells of Perak. Proceedings of the Zoological Society of London 59 (3): 330 - 348, DOI: https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="59" year="1891">
von 
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1891
</bibRefCitation>
: 345. 
<bibRefCitation author="Sykes, ER" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="194 - 199" refId="B160" refString="Sykes, ER, 1903. On the land operculate mollusks collected during the &quot;Skeat Expedition&quot; to the Malay peninsula in 1899-1900. Proceedings of the Zoological Society of London 1903: 194 - 199" title="On the land operculate mollusks collected during the &quot; Skeat Expedition &quot; to the Malay peninsula in 1899 - 1900." volume="1903" year="1903">Sykes 1903</bibRefCitation>
: 197, Jalor [Yala Province, Thailand]. 
<bibRefCitation author="van Benthem Jutting, WSS" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="1" pagination="50 - 77" refId="B180" refString="van Benthem Jutting, WSS, 1949. On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells. Bulletin of the Raffles Museum 19: 50 - 77" title="On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells." volume="19" year="1949">van Benthem Jutting 1949</bibRefCitation>
: 57, Gunong Pulai, Johore [Johor State, Malaysia]. 
<bibRefCitation author="van Benthem Jutting, WSS" journalOrPublisher="Basteria" pageId="0" pageNumber="1" pagination="10 - 20" refId="B181" refString="van Benthem Jutting, WSS, 1960. Some notes on land and freshwater Mollusca of Malaya. Basteria 24: 10 - 20" title="Some notes on land and freshwater Mollusca of Malaya." volume="24" year="1960">van Benthem Jutting 1960</bibRefCitation>
: 13, limestone hill near kampong Tebing Tinggi, N. of Kangar, Perlis [Malaysia]. 
<bibRefCitation author="Chan, S-Y" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" pagination="4 - 5" refId="B17" refString="Chan, S-Y, 1998a. A brief collecting trip to Perak, West Malaysia, part one. The Papustyla 12: 4 - 5" title="A brief collecting trip to Perak, West Malaysia, part one." volume="12" year="1998 a">Chan 1998a</bibRefCitation>
: 4, Ipoh, Perak. 
<bibRefCitation author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
: 40, 41. 
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
: 88. 
<bibRefCitation author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
: fig. 5-13f.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Stoliczka" authorityYear="1872" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola" subGenus="Tylotoechus">Pupina (Tylotoechus) aureola</taxonomicName>
- 
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
: 307. 
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
: 34. 
<bibRefCitation author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie" pageId="0" pageNumber="1" pagination="35 - 70" refId="B66" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
: 39.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Mollendorff" authorityYear="1891" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina arula subsp. perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="Arula" subSpecies="perakensis">Pupina arula perakensis</taxonomicName>
[non 
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
]- 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B48" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. 2017</bibRefCitation>
: 40, fig. 15c, Ipoh, Perak.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Vignard" authorityYear="1829" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pupina</taxonomicName>
sp.- 
<bibRefCitation author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
: fig. 5-11b.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Possible syntype</emphasis>
NHMUK 1988.12.4.101 (Figs 
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 &quot; forma minor &quot; from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B)." figureDoi="10.3897/zookeys.1119.85400.figure34" httpUri="https://binary.pensoft.net/fig/735395" pageId="0" pageNumber="1">34F</figureCitation>
, 
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36A</figureCitation>
) from Pinang.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1998-04-04" location="Phra Kayang Cave" specimenCount="6" stateProvince="Kra Buri District, Ranong Province">
CUMZ 12112 (2 shells and 
<specimenCount type="generic">6 specimens</specimenCount>
in ethanol) from 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:820F10D269C8682B14CDE1E8FB5A0A28" name="Phra Kayang Cave" stateProvince="Kra Buri District, Ranong Province">Phra Kayang Cave</location>
, 
<collectingRegion isEnumeration="true">Kra Buri District, Ranong Province</collectingRegion>
, 
<collectingDate value="1998-04-04">4 Apr. 1998</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2007-03-04" collectorName="Na Mueang Waterfall" location="Ko Samui District, Surat Thani Province" specimenCount="3" stateProvince="Ko Samui District, Surat Thani Province">
CUMZ 12113 (
<specimenCount type="generic">3 specimens</specimenCount>
in ethanol) from 
<collectorName>Na Mueang Waterfall</collectorName>
, 
<collectingRegion isEnumeration="true">Ko Samui District, Surat Thani Province</collectingRegion>
, 
<collectingDate value="2007-03-04">4 Mar. 2007</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2015-12-03" collectorName="Na Mueang Waterfall" location="Ko Samui District, Surat Thani Province" specimenCount="5" stateProvince="Ko Samui District, Surat Thani Province">
CUMZ 12114 (
<specimenCount type="generic">5 specimens</specimenCount>
in ethanol) from 
<collectorName>Na Mueang Waterfall</collectorName>
, 
<collectingRegion isEnumeration="true">Ko Samui District, Surat Thani Province</collectingRegion>
, 
<collectingDate value="2015-12-03">3 Dec. 2015</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2016-12-06" collectorName="Pra Puttabhat Sri Suratth Temple" location="Kanchanadit District, Surat Thani Province" specimenCount="4" stateProvince="Kanchanadit District, Surat Thani Province">
CUMZ 12115 (
<specimenCount type="generic">4 specimens</specimenCount>
in ethanol) from 
<collectorName>Pra Puttabhat Sri Suratth Temple</collectorName>
, 
<collectingRegion isEnumeration="true">Kanchanadit District, Surat Thani Province</collectingRegion>
, 
<collectingDate value="2016-12-06">6 Dec. 2016</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2017-07-04" collectorName="Fig., Khiri Rat Phatthana Temple" location="Wiang Sa District, Surat Thani Province" specimenCount="7" stateProvince="Wiang Sa District, Surat Thani Province">
CUMZ 12116 (
<specimenCount type="generic">7 specimens</specimenCount>
in ethanol; 
<collectorName>Fig.</collectorName>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Opercula of three mainland Southeast Asian Pupina species groups A Pupina bensoni sp. nov. from the Pupina artata species group, specimen CUMZ 12047 B Pupina siamensis from the Pupina arula species group, specimen CUMZ 12067, and C Pupina aureola from the Pupina aureola species group, specimen CUMZ 12116. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure19" httpUri="https://binary.pensoft.net/fig/735380" pageId="0" pageNumber="1">19C</figureCitation>
) from 
<collectorName>Khiri Rat Phatthana Temple</collectorName>
, 
<collectingRegion isEnumeration="true">Wiang Sa District, Surat Thani Province</collectingRegion>
, 
<collectingDate value="2017-07-04">4 July 2017</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2017-03-11" collectorName="Fig." location="Lod Cave" specimenCount="42" stateProvince="Nopphitam District, Nakhon Si Thammarat District">
CUMZ 12117 (4 shells and 
<specimenCount type="generic">42 specimens</specimenCount>
in ethanol; 
<collectorName>Fig.</collectorName>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">32G</figureCitation>
) from 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:ECB345616F6A2E67869DB84ABD3ECE2D" name="Lod Cave" stateProvince="Nopphitam District, Nakhon Si Thammarat District">Lod Cave</location>
, 
<collectingRegion isEnumeration="true">Nopphitam District, Nakhon Si Thammarat District</collectingRegion>
, 
<collectingDate value="2017-03-11">11 Mar. 2017</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2017-03-11" location="Kaeo Surakan Cave" specimenCount="1" stateProvince="Lan Saka District, Nakhon Si Thammarat Province">
CUMZ 12118 (1 shell) from 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:1E7B4ABB898CCF2FADFDA386E4058BEA" name="Kaeo Surakan Cave" stateProvince="Lan Saka District, Nakhon Si Thammarat Province">Kaeo Surakan Cave</location>
, 
<collectingRegion isEnumeration="true">Lan Saka District, Nakhon Si Thammarat Province</collectingRegion>
, 
<collectingDate value="2017-03-11">11 Mar. 2017</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2003-04-04" collectorName="Tham Thong Panara Temple" location="Tham Phannara District, Nakhon Sri Thammarat Province" specimenCount="6" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">
CUMZ 12119 (
<specimenCount type="generic">6 specimens</specimenCount>
in ethanol) from 
<collectorName>Tham Thong Panara Temple</collectorName>
, 
<collectingRegion isEnumeration="true">Tham Phannara District, Nakhon Sri Thammarat Province</collectingRegion>
, 
<collectingDate value="2003-04-04">4 Apr. 2003</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2006-10-11" collectorName="Tham Thong Panara Temple" location="Tham Phannara District, Nakhon Sri Thammarat Province" specimenCount="1" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">
CUMZ 12120 (36 shells and 
<specimenCount type="generic">1 specimen</specimenCount>
in ethanol) from 
<collectorName>Tham Thong Panara Temple</collectorName>
, 
<collectingRegion isEnumeration="true">Tham Phannara District, Nakhon Sri Thammarat Province</collectingRegion>
, 
<collectingDate value="2006-10-11">11 Oct. 2006</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2012-06-11" collectorName="Figs" location="Tham Thong Panara Temple" specimenCount="100" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">
CUMZ 12121 (&gt; 
<specimenCount type="generic">100 specimens</specimenCount>
in ethanol; 
<collectorName>Figs</collectorName>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">32H</figureCitation>
, 
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36F</figureCitation>
, 
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G)." figureDoi="10.3897/zookeys.1119.85400.figure38" httpUri="https://binary.pensoft.net/fig/735399" pageId="0" pageNumber="1">38A</figureCitation>
) from 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:F22207C5A6FCB89A905B9D770931BF73" name="Tham Thong Panara Temple" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">Tham Thong Panara Temple</location>
, 
<collectingRegion isEnumeration="true">Tham Phannara District, Nakhon Sri Thammarat Province</collectingRegion>
, 
<collectingDate value="2012-06-11">11 June 2012</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2014-01-15" collectorName="Tham Thong Panara Temple" location="Tham Phannara District, Nakhon Sri Thammarat Province" specimenCount="15" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">
CUMZ 12122 (
<specimenCount type="generic">15 specimens</specimenCount>
in ethanol) from 
<collectorName>Tham Thong Panara Temple</collectorName>
, 
<collectingRegion isEnumeration="true">Tham Phannara District, Nakhon Sri Thammarat Province</collectingRegion>
, 
<collectingDate value="2014-01-15">15 Jan. 2014</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2017-07-04" collectorName="Tham Thong Panara Temple" location="Tham Phannara District, Nakhon Sri Thammarat Province" specimenCount="1" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">
CUMZ 12123 (12 shells) from 
<collectorName>Tham Thong Panara Temple</collectorName>
, 
<collectingRegion isEnumeration="true">Tham Phannara District, Nakhon Sri Thammarat Province</collectingRegion>
, 
<collectingDate value="2017-07-04">4 July 2017</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2017-07-05" collectorName="Figs" country="Thailand" location="Talot Cave" specimenCount="1" stateProvince="Thung Song District, Nakhon Sri Thammarat Province">
CUMZ 12124 (3 shells and 
<specimenCount type="generic">1 specimen</specimenCount>
in ethanol; 
<collectorName>Figs</collectorName>
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36B</figureCitation>
, 
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G)." figureDoi="10.3897/zookeys.1119.85400.figure38" httpUri="https://binary.pensoft.net/fig/735399" pageId="0" pageNumber="1">38B</figureCitation>
) from 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:8401E5DA6353DD6B47E252250844E15D" country="Thailand" name="Talot Cave" stateProvince="Thung Song District, Nakhon Sri Thammarat Province">Talot Cave</location>
, 
<collectingRegion isEnumeration="true">Thung Song District, Nakhon Sri Thammarat Province</collectingRegion>
, 
<collectingCountry name="Thailand">Thailand</collectingCountry>
, 
<collectingDate value="2017-07-05">5 July 2017</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2015-08-06" country="Thailand" location="Mueang" municipality="Nam Phut Cave" specimenCount="1" stateProvince="Phang Nga District">
CUMZ 12125 (1 shell and 
<specimenCount type="generic">1 specimen</specimenCount>
in ethanol) from 
<collectingMunicipality>Nam Phut Cave</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:665637C4BD0D6A7E4D6B2FBF1E9CB364" country="Thailand" municipality="Nam Phut Cave" name="Mueang" stateProvince="Phang Nga District">Mueang</location>
<collectingRegion country="Thailand" name="Phangnga">Phang Nga District</collectingRegion>
, 
<collectingRegion country="Thailand" name="Phangnga">Phang Nga Province</collectingRegion>
, 
<collectingDate value="2015-08-06">6 Aug. 2015</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2006-10-06" collectorName="Figs" country="Thailand" location="Huai Yot District" specimenCount="1" stateProvince="Trang Province">
CUMZ 12126 (9 shells; 
<collectorName>Figs</collectorName>
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36C</figureCitation>
, 
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G)." figureDoi="10.3897/zookeys.1119.85400.figure38" httpUri="https://binary.pensoft.net/fig/735399" pageId="0" pageNumber="1">38C</figureCitation>
) from Khao Huai Haeng Temple, 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:119318DDEAAC67C8D1AE6F8E3946EAD4" country="Thailand" name="Huai Yot District" stateProvince="Trang Province">Huai Yot District</location>
, 
<collectingRegion country="Thailand" name="Trang">Trang Province</collectingRegion>
, 
<collectingDate value="2006-10-06">6 Oct. 2006</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2006-10-06" collectorName="Ban Khao Poon" country="Thailand" location="Huai Yot District" specimenCount="5" stateProvince="Trang Province">
CUMZ 12127 (
<specimenCount type="generic">5 specimens</specimenCount>
in ethanol) from 
<collectorName>Ban Khao Poon</collectorName>
, 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:DCE0361EC8520005C9FFFFAE4736AD0E" country="Thailand" name="Huai Yot District" stateProvince="Trang Province">Huai Yot District</location>
, 
<collectingRegion country="Thailand" name="Trang">Trang Province</collectingRegion>
, 
<collectingDate value="2006-10-06">6 Oct. 2006</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="1999-08-06" collectorName="Botanical Garden" country="Thailand" location="Yan Ta Khao District" specimenCount="1" stateProvince="Trang">
CUMZ 12128 (1 shell) from 
<collectingRegion country="Thailand" name="Trang">Trang</collectingRegion>
<collectorName>Botanical Garden</collectorName>
, 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:22762CA77470A4E168F8FFAF2096F1F3" country="Thailand" name="Yan Ta Khao District" stateProvince="Trang">Yan Ta Khao District</location>
, 
<collectingRegion country="Thailand" name="Trang">Trang Province</collectingRegion>
, 
<collectingDate value="1999-08-06">6 Aug. 1999</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2017-07-08" country="Thailand" location="Na Yong District" municipality="Khao Pu Chao Bureau of Monks" specimenCount="4" stateProvince="Trang Province">
CUMZ 12129 (
<specimenCount type="generic">4 specimens</specimenCount>
in ethanol) from 
<collectingMunicipality>Khao Pu Chao Bureau of Monks</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:337ED40940C45624F23EAAE58810BE66" country="Thailand" municipality="Khao Pu Chao Bureau of Monks" name="Na Yong District" stateProvince="Trang Province">Na Yong District</location>
, 
<collectingRegion country="Thailand" name="Trang">Trang Province</collectingRegion>
, 
<collectingDate value="2017-07-08">8 July 2017</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2009-01-15" collectorName="Figs" country="Thailand" location="Khlong Thom District" municipality="Sra Morakot" specimenCount="1" stateProvince="Krabi Province">
CUMZ 12130 (8 shells; 
<collectorName>Figs</collectorName>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Umbilical, columellar and parietal views of A Pupinella mansuyi, specimen CUMZ 12148 from Pha Chu, Nan B Pupina artata from the Pupina artata species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai C Pupina godwinausteni sp. nov. from the Pupina arula species group, holotype CUMZ 12090 / 1 D Pupina aureola from the Pupina aureola species group, specimen CUMZ 12130 from Sra Morakot, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1">10D</figureCitation>
, 
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36D</figureCitation>
, 
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G)." figureDoi="10.3897/zookeys.1119.85400.figure38" httpUri="https://binary.pensoft.net/fig/735399" pageId="0" pageNumber="1">38D</figureCitation>
) from 
<collectingMunicipality>Sra Morakot</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:A59C92B6842316F7FDF3DF52B24C19E6" country="Thailand" municipality="Sra Morakot" name="Khlong Thom District" stateProvince="Krabi Province">Khlong Thom District</location>
, 
<collectingRegion country="Thailand" name="Krabi">Krabi Province</collectingRegion>
, 
<collectingDate value="2009-01-15">15 Jan. 2009</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2012-05-17" collectorName="Sra Morakot" country="Thailand" location="Khlong Thom District" specimenCount="2" stateProvince="Krabi Province">
CUMZ 12131 (
<specimenCount type="generic">2 specimens</specimenCount>
in ethanol) from 
<collectorName>Sra Morakot</collectorName>
, 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:27334F659AFD56B92979B7E4BB2ED61E" country="Thailand" name="Khlong Thom District" stateProvince="Krabi Province">Khlong Thom District</location>
, 
<collectingRegion country="Thailand" name="Krabi">Krabi Province</collectingRegion>
, 
<collectingDate value="2012-05-17">17 May 2012</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2008-04-07" collectorName="Toe Bu Cliff Viewpoint" country="Thailand" location="Mueang" specimenCount="15" stateProvince="Satun District">
CUMZ 12132 (
<specimenCount type="generic">15 specimens</specimenCount>
in ethanol) from 
<collectorName>Toe Bu Cliff Viewpoint</collectorName>
, 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:355E27B8F298F29C2C699FD81ABAA124" country="Thailand" name="Mueang" stateProvince="Satun District">Mueang</location>
<collectingRegion country="Thailand" name="Satun">Satun District</collectingRegion>
, 
<collectingRegion country="Thailand" name="Satun">Satun Province</collectingRegion>
, 
<collectingDate value="2008-04-07">7 Apr. 2008</collectingDate>
</materialsCitation>
. 
<materialsCitation collectingDate="2009-01-13" collectorName="Figs" country="Thailand" location="Thung Wa District" municipality="Khantiphol Cave" specimenCount="1" stateProvince="Satun Province">
CUMZ 12133 (7 shells; 
<collectorName>Figs</collectorName>
36E, 38E) from 
<collectingMunicipality>Khantiphol Cave</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:49E05BF3EA3C148A31465911288B5A40" country="Thailand" municipality="Khantiphol Cave" name="Thung Wa District" stateProvince="Satun Province">Thung Wa District</location>
, 
<collectingRegion country="Thailand" name="Satun">Satun Province</collectingRegion>
, 
<collectingDate value="2009-01-13">13 Jan. 2009</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell ovate to fusiform; last whorl ca. 70-75% of shell height. Apertural lip thickened to highly thickened but not expanded. Parietal tooth thickened, fin-shaped or tooth-like, always located next to but not covering posterior canal; columellar tooth fin-shaped, thickened, located next to anterior canal. Posterior canal slightly bulging outward.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="aureola">
<emphasis italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. stoliczkai" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="stoliczkai">
<emphasis italics="true" pageId="0" pageNumber="1">P. stoliczkai</emphasis>
</taxonomicName>
sp. nov. in shell shape and having both fin-shaped and highly thickened parietal and columellar teeth located next to their respective canals; the posterior canal slightly bulges outward. However, 
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="aureola">
<emphasis italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
</taxonomicName>
does not have a furrow between inner and outer peristomes.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Malaysia and southern Thailand (
<bibRefCitation author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species has high variation in shell shape from ovate to fusiform, and the parietal tooth varies from fin-shaped to tooth-like. Despite those shell variations, we assign these shell morphs to 
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="aureola">
<emphasis italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
</taxonomicName>
due to the uniform position of a parietal tooth that is always located next to the posterior canal, and a columellar tooth that is always fin-shaped and not extending over the apertural lip.
</paragraph>
<paragraph pageId="0" pageNumber="1">
By comparing with the possible type specimen, the specimen of 
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. arula subsp. perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" rank="subSpecies" species="Arula" subSpecies="perakensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. arula perakensis</emphasis>
</taxonomicName>
figured in 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B48" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. (2017</bibRefCitation>
: fig. 15c) from Gunung Datok Plot, Ipoh, Perak should belong to 
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="aureola">
<emphasis italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
</taxonomicName>
(Foon, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>