Records and redescription of a mygalomorph spider genus ignored for over 100 years with a new species: the genus Atmetochilus Simon, 1887 (Araneae Nemesiidae) in Thailand
Author
Kunsete, Chawakorn
0000-0002-1594-3736
Author
Sivayyapram, Varat
0000-0002-8466-0270
Author
Traiyasut, Prapun
Author
Thanoosing, Chawatat
Author
Khamwan, Kitiwat
Author
Warrit, Natapot
text
Zootaxa
2020
2020-07-24
4819
3
521
543
journal article
8982
10.11646/zootaxa.4819.3.5
15bf44e4-c720-42d7-b1ec-ab13da78e2a7
1175-5326
4397082
AFB116F2-098E-4CE0-B3AB-77932E0B28F4
Atmetochilus
Simon, 1887
Atmetochilus
Simon, 1887: 109
;
1892: 109
; 65, fig. 73.
Type
species.
Atmetochilus fossor
Simon, 1887
, by monotypy.
Diagnosis.
Differs from other genera in the subfamily
Bemmerinae Simon, 1903
by having very large irregular posterior sternal sigilla centrally confluent. It is most closely related to
Damarchus
Thorell, 1891
and
Darmarchilus
Siliwal, Molur & Raven, 2015
from which it differs by the large sternal sigilla.
Description.
Fovea short, U-shaped. Eyes on tubercle. ALE are largest eyes. No cuspules on labium. Maxilla: serrula absent;
ca
. 30 cuspules on proximal anterior margin of maxillae in both sexes; in male, maxillary cuspules normally smaller than in female, incipient. Posterior sternal sigilla large and medially confluent with middle pair; anterior sigilla small, indistinct. Palpal tarsus of females with one or two spines; no spines on any leg tarsi of males or females. Males with one sigmoidal (S-shaped) row of teeth on all paired claws; females with 2 rows of teeth on all paired claws; female palp claw with one row of teeth on inner surface of claw. Preening combs normally present on metatarsi III and IV. Cymbium without spines. 4 spinnerets, PMS broad, short and straight, dense small spigots on apical surface and separated by the same distance as their basal diameter. PLS long, triangular, digitiform and length of apical segment as long as median segment, basal segment longest, dense small spigots on ventral surface of all segments. Palpal bulb simple and pyriform; embolus elongate but short in
A. koponeni
,
A. lehtineni
and
A. sumatranus
. Distal tibia I of males with ventral spur and megaspines. Spermathecae divided into two lobes. Head of spermathecae semicircular in
A. songsangchotei
sp. nov.
,
A. fossor
and
A. atriceps
, curve in
A. koponeni
,
A. lehtineni
and
A. sumatranus
.
Remarks.
Atmetochilus
was first placed in the family
Ctenizidae
Thorell, 1887
and only transferred to the family
Nemesiidae
much later (
Raven 1985
).
1)
The
type
of
A. fossor
in ZSI, Indian
Museum is unavailable; not in the Muséum National d’Histoire Naturelle, Paris (
Raven 1985
).
2)
This genus description was adapted from
Raven (1985)
,
Zonstein and Marusik (2016)
and also our observations.
Distribution.
Myanmar
,
India
,
Indonesia
(
Sumatra
) and
Thailand
.
Species included.
A. fossor
Simon, 1887
,
A. atriceps
Pocock, 1900
,
A. koponeni
Zonstein and Marusik, 2016
,
A. lehtineni
Zonstein and Marusik, 2016
and
A. sumatranus
Zonstein and Marusik, 2016
,
Atmetochilus songsangchotei
Kunsete and Warrit
,
sp. nov.