Two new Atemnus species (Pseudoscorpiones: Atemnidae) from Xinjiang, China
Author
Zhao, Lingchen
0000-0001-8477-836X
The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, P. R. China & lingchenzhao @ 126. com; https: // orcid. org / 0000 - 0001 - 8477 - 836 X
Author
Gao, Zhizhong
0000-0002-6666-8746
Department of Biology, Xinzhou Teachers University, Xinzhou 034000, Shanxi Province, P. R. China. gaozhizhong 1987 @ 126. com; https: // orcid. org / 0000 - 0002 - 6666 - 8746
Author
Zhang, Feng
0000-0002-3347-1031
The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, P. R. China & dudu 06042001 @ 163. com; https: // orcid. org / 0000 - 0002 - 3347 - 1031
text
Zootaxa
2020
2020-11-02
4869
2
264
274
journal article
8938
10.11646/zootaxa.4869.2.6
39d1c5b2-8a95-458a-b992-fc89b0a5c800
1175-5326
4443644
B3A61385-F017-4C99-A4DB-DEE222B186E6
Atemnus hamiensis
sp. nov.
(
Figs 1A
,
2–4
)
http://zoobank.org/
50A88499-059E-4F64-96FC-29043A0ADDD6
Type material.
Holotype
male (Ps.-
MHBU-XJ18072801
):
China
,
Xinjiang Uygur
Autonomous Region
,
Hami City
,
Yizhou District
,
Toudaogou Village
[
43.124591°N
,
93.795426°E
],
1345 m
a.s.l.
,
28 July 2018
,
Zhizhong Gao
and
Weitong Wang
leg.
Paratypes
: one female (Ps.-
MHBU-XJ
18072802
), one male (Ps.-
MHBU-XJ
18072804
) and one male (ZMH-A0013066), same data as holotype
.
Etymology.
The specific name refers to the
type
locality, Hami City in
China
.
Diagnosis.
The new species can be distinguished from
A. limuensis
by the slender pedipalpal femur of males (2.57–2.71 times longer than deep
vs
2.22–2.55 times in
A. limuensis
) and by the pedipalpal trochanter with a conical dorsal tubercles (pedipalpal trochanter with two distinct coniform protuberances in
A. limuensis
); from
A. politus
by the stouter femur of females (2.37 times longer than deep
vs
2.67–2.86 times in
A. politus
) and by the slender leg IV of males (tibia 3.81–4.00 times longer than deep, tarsus 4.10–4.50 times
vs
tibia 3.33–3.53 times, tarsus 3.58–4.00 times in
A. politus
); from
A. letourneuxi
by the presence of strong granulation on the prolateral surface of pedipalpal patella and by the slender pedipalpal femur of males (2.57–2.71 times longer than deep
vs
2.20 times in
A. letourneuxi
); from
A. strinatii
by the strong granulation on the pedipalpal femur and patella and having distinct eyespots; from
A. syriacus
by the slender pedipalp of males (femur 2.57–2.71 times longer than deep, patella 2.13–2.57 times, chela with pedicel 2.81–3.10 times, chela without pedicel 2.59–2.83 times
vs
femur 2.1 times, patella 1.7 times, chela with pedicel 2.6 times, chela without pedicel 2.3 times in
A. syriacus
) and by the lack of a brown transverse band on the carapace.
Description.
Male (
Fig. 2A
). Carapace and abdomen brown, trochanter of pedipalp yellow-brown and other part deep reddish-brown, legs and chelicerae light yellow; body setae acuminate and long.
Carapace (
Figs 3A
,
4A
): Smooth, with two distinct eyespots; without transverse furrow; with 38 setae, including 4 on anterior margin and 8 on posterior margin, anterior half darker than the posterior half.
Chelicerae (
Fig. 4B
): Hand with 4 setae,
bs
and
es
dentate apically,
is
and
ls
long and acute; fixed finger with 4–5 retrorse teeth and 2 small apical teeth, movable finger with a long, tooth-like subapical lobe; with two lyrifissures on hand; galea (
Fig. 4C
) relatively long (shorter than female), with 6 branchlets; rallum (
Fig. 4E
) with 4 blades, the distal one anteriorly dentated; serrula exterior with 20–21 lamellae.
Pedipalp (
Figs 3E
,
4F
): The retrolateral surface of trochanter, the prolateral surface of femur, patella and hand granular; trochanter with a conical dorsal tubercle; fixed finger (
Fig. 4I
) with about 24 marginal teeth, movable finger with about 28 marginal teeth; trochanter 1.80–1.95, femur 2.57–2.71, patella 2.13–2.42, chela (with pedicel) 2.81–3.10, chela (without pedicel) 2.59–2.83 times longer than deep, movable finger 0.78–0.82 times longer than hand (without pedicel). Venom duct curved and short, nodus ramosus medially between trichobothria
et
and
est
.
st
closer to
sb
than to
t, t
in the middle of movable finger; on fixed chelal finger,
est
midway between
esb
and
eb
, distance between
est
and
esb
nearly equal to that of
ist
and
isb
, distance between
it
and fingertip nearly equal to distance between
ist
and
isb.
Abdomen: Tergites I–IV and XI undivided and other tergites incompletely divided; each tergite and sternite with 2 lyrifissures; tergal chaetotaxy (I–XI): 9: 8: 8: 9: 5–5: 5–4: 5–5: 5–5: 6–5: 6–6 (8 + 4T): 10 + 2T; sternites XI undivided and other sternites divided, sternal chaetotaxy (IV–XI): 5–5: 6–5: 6–7: 6–6: 6–7: 7–6: 7–7 (10 + 4T): 8 + 4T; anterior genital operculum (
Figs 3C
,
4J
) with 8–10 setae on each side; posterior genital sternite with 10 setae; male genitalia (
Fig. 4K
): the distal part (l) of lateral apodemes relatively small, and the hooked branch (b) bowed distally and terminated in a plate-like tip, the proximal part with a distinct dark sclerotized bar (c); the longitudinal fold of medial diverticula (d) with a projection midway along its length; the ejaculatory canal atrium (e) not well-developed, curved distally; the lateral rods (f) long and diverging proximally; the tip of dorsal apodeme (g) completely joined; the ventral diverticulum (h) bilobed.
FIGURE 1.
Live images of
Atemnus hamiensis
sp. nov.
(A) and
Atemnus niger
sp. nov.
(B); C, D. Collecting sites of
Atemnus niger
sp. nov.
FIGURE 2.
Atemnus hamiensis
sp. nov.
, A. Holotype male, dorsal view; B. Paratype female, dorsal view.
Legs: Leg I (
Figs 3D
,
4G
) and leg IV (
Figs 3H
,
4H
) typical. Fairly smooth, claws simple, longer than arolium, subterminal seta acute and curved. Leg IV with one tactile seta at the base of tarsus (
Fig. 3G
) (TS = 0.14–0.17).
Female (
paratype
) (
Fig. 2B
): Mostly same as the male
holotype
.
Chelicerae. Hand with 4 setae; galea (
Fig. 4D
) long with 6 branchlets; serrula exterior with 20 lamellae.
Pedipalps. Fixed finger with about 22 marginal teeth, movable finger with about 27 marginal teeth; trochanter 1.77, femur 2.37, patella 2.16, chela (with pedicel) 2.73, chela (without pedicel) 2.46 times longer than deep, movable finger 0.80 times longer than hand (without pedicel).
Abdomen. Tergites I–II and XI undivided and other tergites incompletely divided; each tergite and sternite with 2 lyrifissures; tergal chaetotaxy (I–XI): 7: 7: 4–4: 5–5: 5–5: 5–5: 5–5: 7–5: 5–5: 7–6(9 + 4T): 10 + 2T; sternites XI undivided and other sternites divided, sternal chaetotaxy (IV–XI): 5–5: 8–6: 6–6: 5–6: 5–5: 7–7: 7–7 (10 + 4T): 8 + 4T; genital area (
Figs 3F
,
4L
) with 9 setae on the anterior genital operculum and posterior margin; genital structure (
Fig. 4M
) simple, spermathecae provided with separated median cribriform plates.
Dimensions (length/breadth or depth, in mm). Males. Body length 3.14–3.16. Carapace 0.72–0.84/0.53–0.66 (1.27–1.36). Pedipalp: trochanter 0.39–0.45/0.20–0.25 (1.80–1.95), femur 0.65–0.77/0.24–0.30 (2.57–2.71), patella 0.64–0.80/0.30–0.33 (2.13–2.42), chela (with pedicel) 1.04–1.24/0.37–0.40 (2.81–3.10), chela (without pedicel) 0.96–1.13 (2.59–2.83), hand length (without pedicel) 0.55–0.65 (1.49–1.63), movable finger length 0.45–0.51 (0.78–0.82 times longer than length of hand without pedicel). Leg I: femur 0.21–0.25/0.15–0.19 (1.32–1.40), patella 0.35–0.39/0.14–0.17 (2.29–2.50), tibia 0.36–0.43/0.10–0.12 (3.58–3.60), tarsus 0.31–0.38/0.07–0.09 (4.22–4.43). Leg IV: femur + patella 0.64–0.76/0.22–0.27 (2.81–2.91), tibia 0.52–0.61/0.13–0.16 (3.81–4.00), tarsus 0.36–0.41/0.08–0.10 (4.10–4.50).
Female. Body length 3.95. Carapace 0.84/0.59 (1.42). Pedipalp: trochanter 0.39/0.22 (1.77), femur 0.64/0.27 (2.37), patella 0.67/0.31 (2.16), chela (with pedicel) 1.12/0.41 (2.73), chela (without pedicel) 1.01 (2.46), hand (without pedicel) 0.56 (1.37), movable finger length 0.45 (0.80 times longer than hand without pedicel). Leg I: femur 0.21/0.12 (1.75), patella 0.36/0.11 (3.27), tibia 0.34/0.10 (3.4), tarsus 0.33/0.07 (4.71). Leg IV: femur + patella 0.69/0.21 (3.29), tibia 0.54/0.14 (3.86), tarsus 0.34/0.09 (3.78).
Distribution.
Known only from the
type
locality.
Habitat.
All specimens were collected under the tree bark of
Populus
(Salicaceae)
, the specific species name unknown.