Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia Author Kovařík, František Author Šťáhlavský, František text Euscorpius 2020 315 1 37 journal article 6637 10.5281/zenodo.4648646 9797e168-d1b1-4c0a-8786-638b6f992159 1536-9307 4648646 CD6F50BE-BFE9-49A9-BAA2-4C6C2B20C7E9 Euscorpius sadileki sp . n . ( Figures 115–149 , 184 , 186 , Tables 3–4 ) http : //zoobank . org / urn : lsid : zoobank . org : act : 382FE1BA- 8307-406D-A493-6EC930BAA309 TYPE LOCALITY AND TYPE REPOSITORY . Serbia , Sokobanja , 43.63°N 21.80°E ; FKCP . TYPE MATERIAL . Serbia , Sokobanja , 43.63°N 21.80°E , 14. VI .2014, leg. D. SadÍlek, 1♂ 1♀ ( holotype and paratype , 543), FKCP ; Nišava Province , Gadžin Han env., road between Donji Dušnik and Supotnica, Suva Planina foothills, 43.155ºN 22.118ºE , 5. VI .2014, 1♀ ( paratype ), leg. P. ŠÍpek et al., FKCP . ETYMOLOGY. The species epithet is a patronym honoring David SadÍlek, the collector of holotype of the new species. DIAGNOSIS ( ♂ ♀ ). Total length 26–36 mm . Color reddish black, legs and telson yellow. Pedipalp patellar external trichobothria numbers: 4 eb , 4 eb a , 2 esb , 4 em , 4 est , 6 et , ventral aspect of patella with 8–9 trichobothria. Pectinal teeth number 9 in male, 6–8 in females. Chelicerae yellow, with reticulation. Male with pedipalp fingers type C, female with fingers undulate. Dorsal metasomal carinae on III–IV segments granulated. Dorsolateral metasomal carinae on II–IV segments absent. Ventrolateral metasomal carinae on II–IV segments present and smooth or granulate (mainly in male). Metasoma II–IV densely (roughly) granulated including lateral surface. Metasoma V ventrally granulated with median carina present. Chela length/width ratio, 2.2 in male and 2.4–2.5 in females. Metasoma IV length/ width ratio, 1.68 in male and 1.8 in females. Telson length/ depth ratio, 2.6 in male and 3–3.1 in females. DESCRIPTION ( ♀ ♂ ). Total length about 26–36 mm in both sexes. The habitus is shown in Figs. 115–118 . For position and distribution of trichobothria of pedipalps, see Figs. 141–147 . Sexual dimorphism, see below in the description. Coloration ( Figs. 115–118 , 149 ). Base color uniformly reddish black, sternites lighter, yellowish brown, legs and telson yellow. Chelicerae yellow, reticulate. Carapace and mesosoma ( Figs. 115–122 ). Carapace finely granulated with several smooth areas; carinae are absent. The anterior margin of the carapace is straight. Carapace with two lateral eyes. Tergites finely granulated, without developed carina. Tergite VII lacking lateral and median carinal pairs. Sternites III–VII rather smooth in females and sparsely finely granulated in male, both lacking lateral and median carinae. Stigmata small, narrow ellipical. Pectinal teeth number 9 in male and 6–8 (1 x 6, 1 x 7, 2 x 8) in females, fulcra are present. Pectines have three marginal lamellae and 4–6 middle lamellae. Figures 115–118 : E. sadileki sp . n . Figures 115–116 . Male holotype in dorsal (115) and ventral (116) views. Figures 117–118 . Female paratopotype in dorsal (117) and ventral (118) views. Scale bar: 10 mm. Figures 119–125 : E. sadileki sp . n . Figures 119 , 121 , 123–125 . Male holotype, carapace and tergites I–IV (119), coxosternal area and sternites (121), left legs II–IV, retrolateral aspect (123–125 respectively). Figures 120 , 122 . Female paratopotype, carapace and tergites I–III (120), coxosternal area and sternites III–IV (122). Figures 126–133 : E. sadileki sp . n . Figures 126–129 . Female paratopotype, telson lateral (126), and metasoma and telson lateral (127), dorsal (128), and ventral (129) views. Figures 130–133 . Male holotype, telson lateral (130), and metasoma and telson lateral (131), dorsal (132), and ventral (133) views. Scale bars: 5 mm (127–129, 131–133). Figures 134–140 . E. sadileki sp . n . , male holotype, pedipalp segments. Chela dorsal (134), external (135) and ventral (136) views. Patella dorsal (137), external (138) and ventral (139) views. Trochanter and femur dorsal (140) view. Figures 141–148 : E. sadileki sp . n . , female paratopotype, pedipalp segments. Chela dorsal (141), external (142) and ventral (143) views. Patella dorsal (144), external (145) and ventral (146) views. Trochanter and femur dorsal (147) and ventral (148) views. Trichobothrial pattern is indicated by white circles. Figure 149 . E. sadileki sp . n . , female paratype in vivo habitus. Metasoma and telson ( Figs. 126–133 ). Metasoma very sparsely hirsute. Metasoma I–V granulated finely in females and roughly in male. Dorsal metasomal carinae on I–V segments distinctively irregularly granulated; dorsolateral carinae absent; ventrolateral carinae present, smooth in females and granulated in male. Metasoma V completely granulated in both sexes with median carina developed, on metasoma I–IV ventral median carina is absent. Anal arch with small pigmented granules. Telson finely granulated, elongate in females and swollen in males, with annular ring present in both sexes. Aculeus short more curved in male. Pedipalps ( Figs. 134–148 ). Pedipalps very sparsely hirsute. Patella with 24 (4 eb , 4 eb a, 2 esb , 4 em , 4 est , 6 et ) external and 8–9 (3 x 8, 1 x 9) ventral trichobothria. Chela with 4 ventral trichobothria, of which V 4 is located on external surface near Eb 1 . Female paratype has additional trichobothrium eb a on chela external surface ( Fig. 142 ), which is absent in male holotype . Femur and patella are granulated, femur roughly and patella finely. Femur has granulated developed carinae; ventroexternal carina is incomplete to missing. Patella with 5 complete carinae including irregular wide externomedian. DPS weak to moderate. Manus dorsally with fine, rounded granules, which do not form median carina; only five chelal carinae are developed. Male with pedipalp fingers type C, weakly undulate, similar ar females. Legs ( Figs. 123–125 ). Both pedal spurs present on all legs, lacking spinelets; tibial spurs absent. Tarsus with single row of spinules on ventral surface, terminating distally with two essentially adjacent spinules. Measurements . See Table 3 . AFFINITIES. The combination of four characters (pedipalp patellar external trichobothria numbers: 4 eb , 4 eb a ; metasoma IV length/width ratio 1.6–1.8; male with pedipalp fingers type C and female with fingers undulate; and metasoma II– IV densely (roughly) granulated including lateral surface) is unique in the entire genus Euscorpius . DISTRIBUTION. Serbia ( Fig. 184 ).