Faunistic and diagnostic notes on some species of the genus Omalium Gravenhorst, 1802 of Caucasus, Turkey and Iran (Coleoptera: Staphylinidae: Omaliinae, Omaliini) Author Shavrin, Alexey V. 0000-0001-6487-4530 Daugavpils University, Institute of Life Sciences and Technologies, Coleopterological Research Centre, Vienibas 13, Daugavpils, LV- 5401, Latvia. ashavrin @ hotmail. com; https: // orcid. org / 0000 - 0001 - 6487 - 4530 ashavrin@hotmail.com Author Khachikov, Eduard A. Southern Federal University, Bolshaya Sadovaya 105 / 42, Rostov-on-Don, 34006, Russia. text Zootaxa 2023 2023-01-09 5227 4 443 458 journal article 222448 10.11646/zootaxa.5227.4.3 ea087173-f3d9-491b-8157-366b194bb519 1175-5326 7518924 B28C27C8-7273-4294-8487-150075ABD889 Omalium rivulare ( Paykull, 1789 ) ( Figs. 7 , 25–26 , 39 ) Staphylinus rivularis Paykull, 1789: 65 Material examined. RUSSIA : ROSTOV AReA: 1 ♀ : Rostov-on-Don. 26.10.1981 . E.A. Khachikov leg. (cKh); 1 ♀ : same data. 20.05.1982 (cKh); 1 ♀ : same data. 23.11.1991 (cKh); 1 ♁: same data. 09.10.1992 (cKh); 1 ♁: same data. 05.1994 (cKh); 1 ♁, 1 ♀ : same data. 04.1996 (cKh); 1 ♀ : same data. 07.11.2010 (cKh); 1 ♀ : same data, Simferopolskaya str. 13.10.1979 (cKh); 1 ♀ : Shchepkinsk. Forest. 13.04.1991 . E.A. Khachikov leg. (cKh); 2 ♁♁: 10 km NW Gukovo, “Lesnaya Skazka”. 01.06.2000 . E.A. Khachikov leg. (cKh); KRASNODAR TeRRITORY: 1 ♀ : Derbiy, near Ubinskaya. 03.06.1986 . I.A. Ushakov leg. ( ZMM ); 1 ♀ : 20 km Novorossiysk, environs of Malyi Utrish. In cow manure. 10.05.1995 . M. Skomorokhov leg. (cKh); 1 ♁: Temnolesskaya St. 04.10.1999 . V.V. Grebennikov leg. (cKh); STAVROPOL TeRRITORY: 1 ♁: Kislovodsk. 06.1994. E.A.Khachikov leg. (cKh); 2 ♁♁, 2 ♀♀ : Pyatigorsk, Mineralnye Vody, Beshtau Mt. 04– 17.07.1993 . E.A. Khachikov leg. (cKh); 2 ♁♁: same data. 06.1994. E.A. Khachikov leg. (cKh); 1 ♁: Pyatigorsk. 05.1993. E.A. Khachikov leg. (cKh); ADYGeA : 1 ♀ : Caucasus Nature Reserve, W of Guzeripl, Abago Pasture. 1750 m a.s.l., Fagus forest with Abies , Sorbus etc. near the upper border of forest. 24.05.1985 . A.B. Ryvkin leg. (cR); 2 ♁♁, 1 ♀ : Abago, 11 km of Guseripl-Abago road. 05.10.2015 . E.A. Khachikov leg. (cSh); 3 ♁♁, 1 ♀♀ : Maykop, Khanskaya, right bank of Belaya R., 44°41′03.54′′N 39°55′41.25′′E . Flood-plain forest. 15.05.2014 . E.A. Khachikov leg. (cKh, cSh); 1 ♁: Khanskaya. 16.05.2015 . E.A. Khachikov leg. (cKh); 1 ♀ : Maykop District, Kuzhorskaya, territory of Kuzhorskiy Botanic Preserve, 41°41′02.86′′N 40°14′38.93′′N 04.2014. Hornbeam-oak forest. V.A. Brinikh & E.A. Khachikov leg. (cKh); KARACHAY-CHeRKeSSIA : 1 ♀ : Teberda State Nature Biosphere Reserve, Malaya Khotipara Mt. 2600–2800 m a.s.l., subalpic zone. 26.07.2007 . A.S. Prosvirov leg. (cSh); 2 ♁♁: same Reserve, Alibek Gorge, Dvuyazychniy Glacier. 12.07.1996 . E.A. Khachikov leg. (cSh); 1 ♀ : same Reserve, Shumka tract. 18.06.1997 . Yu.G. Arzanov leg. (cSh); 1 ♀ : Teberda. 25.06.1940 . S. Nikulin leg. ( ZIN ); KABARDINO-BALKARIA : 1 ♀ : Adyl-Su near Tyrnyauz. 1900 m a.s.l. 27.06.1991 . J. Martens & W. Schawaller leg. ( SMNS ); DAGeSTAN : 1 ♀ : valley of Samur River, Garakh ca. 35 km SW of Magaramkent. 700–850 m a.s.l., Crataegus , Quercus , Acer , Rosa etc. in drift and litter. 23.10.1987 . S. Golovatch leg. (cSh); 1♀ : Tabasaranskiy District, above Dyubek. 10– 14.05.2008 . E.A. Khachikov leg. (cKh); 1 ♁: Kurush. 1850 m a.s.l. J. Martens & W. Schawaller leg. ( SMNS ); GEORGIA : MTSKHeTA-MTIANeTI : 5 ♁♁, 5 ♀♀ : Saguramo Reserve, NE of Etsketa, Zedacni, 1100–1200 m a.s.l. Forest with Fagus , Carpinus , Acer etc., in litter and under bark. 20.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM ); KVeMO-KARTLI: 1 ♁, 3 ♀♀ : Algeti National Park, W of Manglisi. 1400–1500 m a.s.l., forest with Fagus , Picea , Abies etc., in litter and under bark. 16– 18.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. (cSh); TBILISI : 3 ♁♁, 3 ♀♀ : Tskhneti. 1150–1300 m a.s.l. Forest with Fagus , Carpinus , Acer etc., in litter. 16, 18.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. (cSh, cR); 1 ♁: pass between Tbilisi and Bikhura. 1050 m a.s.l., in litter and soil. 10.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. (cSh); KAKHeTI : 3 ♁♁, 3 ♀♀ : N Kvareli. 700–750 m a.s.l., forest with Fagus , Carpinus , Quercus etc., in litter and under bark. 04– 05.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. (cSh); 3 ♀♀ : Mariamjvari Nature Reserve, NNE of Sagarejo, 1150–1250 m a.s.l., forest with Fagus , Carpinus , Acer , Pinus etc. 13– 14.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. ( ZMM ); ARMENIA : LORI : 7 ♁♁, 4 ♀♀ : Odzun W of Alaverdi. 1500–1550 m a.s.l., forest with Quercus , Fagus , Carpinus etc., in litter and under stones. 23– 24.07.1987 . S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM ); 1 ♁, 1 ♀ : Stepanavan, 1500–1600 m a.s.l., Quercus , Fagus , Carpinus etc. forest, litter and under bark. 21– 22.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. ( ZMM ); ARAGATSOTN : 1 ♁: Idjeva District, Tsakhkevank. 850–900 m a.s.l., forest with Quercus , Acer , Carpinus etc., in litter and tree holes. 25.07.1987 . S.I. Golovatch & K.Yu. Eskov leg. (cSh); TAVUSH : 2 ♁♁, 1 ♀ : Dilijan National Park, Akhnabat Valley near Salakh. 1400–1500 m a.s.l., forest with Fagus , Taxus , Carpinus , etc., in litter, under bark and stones. 17.04.1983 . S.I. Golovatch leg. (cSh); AZERBAIJAN : SHAKI-ZAQATALA: 7 ♁♁, 2 ♀♀ : N of Vartashen. 900 m a.s.l., forest with Fagus and Quercus , in litter. 02.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM ); 1 ♀ : NW above Bashlayski ca. 20 km NNW of Sheki. 1250 m a.s.l., forest with Fagus , Carpinus , Acer etc., in litter. 03.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. (cSh); 1 ♀ : 5 km N of Kutkashen (=Gabala). 1150–1200 m a.s.l., forest with Fagus and Carpinus , in litter. 02.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. ( ZMM ); LANKARAN : 1 ♀ : Chilisa, ca. 7 km N of Kelbadjar. 1400–1500 m a.s.l., forest with Quercus , Carpinus , Acer etc., in litter and under bark. 31.05.1987 . S.I. Golovatch & K.Yu. Eskov leg. ( ZMM ); GEORGIA : ADZHARIA: 1 ♁: W PART Of ADZHARO-IMERETINSKIY MOUNTAIN RANGE,NEAR KLIMO,LEfT BANK Of KINTRISHI RIVER, 41°43’58.20”N 42°04’59.00”E . 11.04- 06.06.2016 . D. FOMINYKH LEG.( CKH ); TURKEY : ISTANBUL : 1 ♁:Yalova-Orhangazi. 11.05.1976 . C. Besuchet leg. ( MHNG ); ZONGULDAK : 2 ♁♁: S Zonguldak. 500 m a.s.l. 23.05.1976 . C. Besuchet & I. Löbl leg. ( MHNG ); KASTAMONU : 1 ♀ : Küre â 5 km N. 600 m a.s.l. 18.05.1976 . C. Besuchet & I. Löbl leg. ( MHNG ); KONYA : 1 ♁, 2 ♀♀ : Sertavul Gegidi. 1500–1600 m a.s.l. 18.04.1978 . C. Besuchet & I. Löbl leg. ( MHNG ); ARTVIN : 1 ♁: Artvin. 800 m a.s.l. 07.06.1986 . C. Besuchet, I. Löbl & D. Burckhardt leg. ( MHNG ); 1 ♁, 1 ♀ : same data. 1500 m a.s.l. 09.06.1986 ( MHNG ); IRAN : MAZANDARAN : 1 ♁: SSO Gol-e-Loveh [Kanaskuh], 37°18′N 55°43′E . 1200 m a.s.l. 21.08.1975 . A. Senglet leg. ( MHNG ); TeHRAN : 1 ♀ : Kandovan, 36°10′N 51°19′E . 12.07.1973 . A. Senglet leg. ( MHNG ). Published records. ARMENIA : Assing & Schülke 2019: 100 ; AZERBAIJAN : Hochhuth 1849: 204 ; GEORGIA: Jakobson 1909: 455; NAGORNO-KARABAKH: Assing & Schülke 2019: 100 ; RUSSIA : ADYGeA : Zanetti 2002: 56 ; DAGeSTAN : Khachikov et al . 2021: 250 ; KARACHAY-CHeRKeSSIA : Khachikov 1998: 6 ; KRASNODAR TeRRITORY: Zanetti 2002: 56 ; NORTH OSSeTIA-ALANIA : Zanetti 2002: 56 ; STAVROPOL TeRRITORY: Khachikov 1998: 6 ; TURKEY : Zanetti 2002: 56 , Bordoni 2010: 49 , Assing 2013: 106 , 2014: 1135 , Anlaş 2019: 217. Distribution. Omalium rivulare is widely distributed species in the Holarctic Region ( Herman 2001 , Schülke & Smetana 2015 ). Remarks. Habitus as in Fig. 7 . Aedeagus as in Figs. 25–26 . Female genital segment as in Fig. 39 . It is here recorded from Iran for the first time.