Phylogeny and systematics of the giant rhinoceros beetles (Scarabaeidae: Dynastini) Author Rowland, J. Mark Author Miller, Kelly B. text Insecta Mundi 2012 2012-11-30 2012 263 1 15 journal article 10.5281/zenodo.5175646 1942-1354 5175646 Tribe DYNASTINI MacLeay Description. Endrödi (1985) constitutes the most recent comprehensive taxonomic review of the Dynastinae , in which he provides the following morphological description of the Dynastini relative to the other tribes of the subfamily: “This tribe includes the biggest species of the whole family Melolonthidae . Most species display very strong sexual dimorphism. Horns of males often very long, head and pronotum [horns] in females usually almost absent. Fore legs of males mostly longer than in females. Also elytra in many species different: in male smooth, in female strongly sculptured. Mandibles mostly incised on apex, outer side straight or lobed. Antennae 10-jointed, club in both sexes short. Form of prosternal process highly variable. Propygidium either with or without a stridulatory area. Claw-joint of anterior tarsi strongly thickened only in Dynastes neptunus Quensel and D. satanas Moser ; apex of basal-joint of hind tarsi rarely spiniformly produced.”