Phylogenetic and morphological resolution of the Helobdella stagnalis species-complex (Annelida: Clitellata: Hirudinea) Author Saglam, Naim Author Kutschera, Ulrich Author Saunders, Ralph Author Saidel, William M. Author Balombini, Katherine L. W. Author Shain, Daniel H. text Zootaxa 2018 2018-04-03 4403 1 61 86 journal article 30344 10.11646/zootaxa.4403.1.3 82592d82-9f4b-4f22-8c93-ca00c436d13b 1175-5326 1212238 7B575B1A-9158-46CD-B275-5A3224693663 Helobdella serendipitious n. sp. ( Figs. 2C – 7C ) ( Holotype : ANSP GI 19495) (LSID: urn:lsid:zoobank.org:act:D2A31CAB-F69D-413A-AD52-1497EB146753 )— Adult 11.2–20.3 mm long, 4.5–5.2 mm wide, width of anterior sucker 1.2–2.4 mm , width of posterior sucker 2.0– 3.6 mm . Dorsal body light to yellowish-grey, sometimes with greenish tint. Ventral color containing mixed translucent creamy and browngreenish spots. Total of 66 annuli; I, II, III uniannulate, IV, V biannulate, VI-XXIII triannulate, XXIV biannulate XXV-XXVII uniannulate. One pair of large circular eyes clearly separated on segment III ( Fig. 7C ). Gonopores located between XI a1/a2 (23th/24th annulus) and XI a2/a3 (24th/25th annulus), respectively. Clitellum on 11th segment. Six pairs of ovoidal, irregular and amorphous testisacs inter-segmentally arranged on XII/XIII–XVII/ XVIII. Ejaculatory bulbs moderately longer, elongated ellipsoid, lying at much lower level by the sides of the atrium, connected by slender ejaculatory ducts with sharp turn backwards into atrium in XI. Ovisacs large, connected with long and curled common oviduct. Length of ovisacs and oviduct almost equal. Ovisac and oviduct 2/3 the length of vas deferens ( Figs. 2C , 4C ). Crop caeca comprising six unbranched pairs. Four pairs thick intestinal caecum curved downward in first and fourth pairs. Rectum short, “S” shaped and similar to that of H. stagnalis . ( Figs. 3C , 5C ).