The Spider Subfamily Amaurobioidinae (Araneae, Anyphaenidae): A Phylogenetic Revision At The Generic Level
Author
RAMÍREZ, MARTÍN J.
text
Bulletin of the American Museum of Natural History
2003
2003-08-13
2003
277
1
262
http://www.bioone.org/doi/abs/10.1206/0003-0090%282003%29277%3C0001%3ATSSAAA%3E2.0.CO%3B2
journal article
10.1206/0003-0090(2003)277<0001:TSSAAA>2.0.CO;2
0003-0090
12776077
Sanogasta puma
,
new species
Figures 76E
,
86E–H
,
87
TYPES
: Female
holotype
and male
paratype
from
Argentina
,
Buenos Aires province
,
Isla
Talavera,
2 km
E Zárate, ca. 34°06̍S, 59°02̍W,
3.XI.1996
, M. Ramírez, deposited in MACNAr 9815.
ETYMOLOGY
: The specific name is a noun in apposition taken from the felid
Puma
concolor
, referring to the uniformly grayish brown color of the body.
DIAGNOSIS
: Resembles
S. tenuis
in having an elongate body without pattern, but can be distinguished by having ovate, voluminous spermathecae, with pointed anterior ends, and a relatively large secondary conductor.
FEMALE
(
holotype
): Total length 6.12. Carapace flattened, without thoracic groove, length 2.47, width 1.50, wider at leg II. Length of tibia/metatarsus: I, 1.35/1.13; II, 1.18/0.98; III, 0.83/0.82; IV, 1.55/1.22. Palpal tarsus length 0.60. Chelicerae unmodified, with two teeth on retromargin. Sternum length 1.57, width 0.82. Spines: leg
I,
femur d 1–1–1, p 0 d1–2, r d1ap; tibia v 2–2–2; metatarsus v 2bas.
II
= I.
III,
femur d 1–1– 1, p and r 0 d1 d1; tibia v p1–2–2, p v1 v1 + 0–1–0 bristle, r d1–1, d r1–0–1 bristles; metatarsus v 2–0–2, p 1–1, r 1–1, d 0 (p1 bristle) 2.
IV,
femur d 1–1–1, p and r d1ap; tibia v p1–2–2, p v1 v1–0 + 0–1–0 bristle, r 1 d1–1, d r1–0–1 bristles; metatarsus v 2– 2–2 or 2 p1–2, p 0 v1 0 v1, r d1–1–1, d 0 (p1 bristle) 0–2 (the p1ap is a bristle). Prolateral spines on tibiae III and IV ventrally displaced. Abdomen length 3.65, width 1.47, spiracle–epigastrium 1.90, spiracle–spinnerets 0.73. Color: grayish brown uniform (fig. 76E), paler on venter, abdomen darkening posteriorly, membranous base of anterior spinnerets dark. Epigyne (figs. 86G, H, 87): anterior pouch advanced, longitudinal, without definite cavity. Copulatory ducts arising contiguously. Ducts of accessory bulbs very short, ventral. Spermathecae ovate, anteriorly acute.
Fig. 86.
A–D.
Sanogasta minuta
(Keyserling)
(Córdoba, Huerta Grande).
A.
Male copulatory bulb, retrolateral view.
B.
Same, ventral view.
C.
Epigyne, ventral view.
D.
Same, cleared.
E–H.
Sanogasta puma
,
n. sp.
E.
Male copulatory bulb, ventral view (Buenos Aires, Punta Lara).
F.
Same, retrolateral view.
G.
Epigyne, ventral view (holotype).
H.
Same, cleared. Scale bars = 0.2 mm.
Fig. 87.
Sanogasta puma
,
n. sp.
, epigyne, ventral view (Buenos Aires, Paraná de Las Palmas).
A.
Anterior pouch.
B.
Copulatory openings.
MALE
(
paratype
): Total length 5.45. Carapace with thoracic groove scarcely marked, length 2.43, width 1.60. Length of tibia/ metatarsus: I, 1.73/1.40; II, 1.35/1.13; III, 0.98/0.95; IV, 1.70/1.37. Chelicerae small, vertical. Sternum length 1.47, width 0.78. Spines as in female, except: leg
I,
tibia p 1 d1 1 0, r 1 0 d1 0, d r1–0–1 bristles; metatarsus p d1.
II,
tibia p 1 d1 1 0, r 1 0 d1 0, d r1–0–1 bristles; metatarsus p d1.
III,
tibia, r 1 d1 0 1 0; metatarsus v 2 p1–2. Abdomen length 2.93, width 1.33, spiracle–epigastrium 1.50, spiracle–spinnerets 0.58. Color as in female. Palp (fig. 86E, F): tibia short, width/ length 1.39, cymbium relatively large. Embolus thin, basal process small, triangular. Median apophysis very thin, short, closely associated with paramedian. Apex of paramedian apophysis short, sinuous, translucent. Secondary conductor not fused to anterior margin of tegulum; canal conspicuous, arising at base of paramedian apophysis; retrolateral portion with basal, conical projection. Anterior margin of tegulum compressed over base of secondary conductor.
NATURAL
HISTORY
: This species lives in the bases of large grasses, mainly pampa grasses (‘‘cortadera’’,
Cortaderia selloana
), in periodically flooded areas. The elongate, flattened body with uniform color and the way the spider aligns the legs with the grass leaves make a strikingly cryptic effect.
DISTRIBUTION
: Southeastern
Brazil
,
Uruguay
, and northeastern
Argentina
.
OTHER
MATERIAL
EXAMINED
:
BRASIL
:
São Paulo
:
Botucatu
,
Fazenda Nossa Senhora da Conceição
, ‘‘parte aérea de
Saccharum officinarum
’’,
4.II.1999
, L.I.
Rinaldi
, B.
Mendez
, 13 (
UNESP
),
20.IV.1999
,
1♀
(
UNESP
)
;
Botucatu
,
Usina São Manoel
,
2.III.1988
, I.
Rinaldi
and L.
Forti
,
1♀
(
UNESP
),
29.IX.1986
, 13 (
UNESP
)
.
URUGUAY
:
Departamento Rocha
:
Arroyo Sarandí del Consejo
, ruta
9 km
251,
18.V.1993
,
M. Ramírez
and
F. Pérez Miles
, 13 (MACNAr)
.
ARGENTINA
:
Santa Fe
:
El Toba
,
4.IV.1968
, M.
Galiano
,
1♀
(
MACNAr
)
.
Entre Ríos
:
Rosario
del
Tala,
XI.1988
, M.
Ramírez
,
1♀
(
MACNAr
, photos MJR 62, 63, 269)
.
Buenos Aires
:
Same
data as types, 13
4♀
1 immature
;
Delta
, estación experimental INTA,
3.II.1983
, M.
Carbajal
and M.
Ramírez
, 13 (
MACNAr
)
;
Delta
,
Paraná de Las Palmas
,
17.VII.1964
, M.
Galiano
,
2♀
(
MACNAr
)
;
Dique Luján
,
26.IX.1982
, P.
Goloboff
and M.
Ramírez
, 13 (
MACNAr
)
;
Mar del Tuyú
, costa atlántica,
2.V.1981
, M.
Ramírez
,
1♀
1 immature
(
MACNAr
)
;
Reserva Otamendi
,
10.VI.1997
, M.
Ramírez
, L.
Compagnucci
, C.
Grismado
, F.
Uehara
,
1 immature
(
MACNAr
)
;
Punta Lara
,
Ensenada
,
10.X.1984
, M.
Galiano
, C.
Scioscia
and P.
Goloboff
, 13 (
MACNAr
),
11.X.1985
,
F. Miranda
and M.
Ramírez
, 13 (
MACNAr
)
;
Río Luján
, estación FCGM, marsh with ‘‘espadaña’’,
5.X.1993
, M.
Ramírez
and A.
Pérez
, 23 (
MACNAr
)
;
Zelaya
,
VII.1938
, J.B.
Daguerre
,
2♀
(
MACNAr
)
.