Lectotypifications, synonymy, and a new name in Capsicum (Solanoideae, Solanaceae)
Author
Barboza, Gloria E.
text
PhytoKeys
2011
2011-02-11
2
23
38
http://dx.doi.org/10.3897/phytokeys.2.730
journal article
http://dx.doi.org/10.3897/phytokeys.2.730
1314-2003-2-23
FFC8FFA07E7426596A663819FFEEFFB0
576065
Capsicum eshbaughii Barboza
nom. nov.
Capsicum eximium
var.
tomentosum
Eshbaugh & P.G.Sm., Baileya 18: 15. 1971, non
Capsicum tomentosum
Kuntze, 1891. - Holotype: BOLIVIA, Dpto. Santa Cruz, Prov. Florida, Mairana area, 1300 m, P.G.Smith Sa281 (holotype, IND!; isotypes: MU! (MU-153648, MU-153649).
Fig. 2
General
Capsicum eximium
var.
tomentosum
was described as an unusual glandular tomentose variety of
Capsicum eximium
Hunz. The presence of this kind of pubescence densely covering the vegetative organs, pedicels, and calyx, and of a 5-10 toothed-calyx (
Fig. 2 N
) in specimens from a restricted area in south-central Bolivia (Dpto. Santa Cruz and Cochabamba) makes them quite different from
Capsicum eximium
. In fact,
Capsicum eximium
has non- glandular pubescence (
Fig. 3 M
), a calyx with only 5 teeth (
Fig. 3 C, E
), and is distributed in a larger area (Bolivia: La Paz to Tarija and Argentina: Jujuy to
Tucuman
).
Figure 2.
Capsicum eshbaughii
.
A
embryo
B, H, F, J, L
glandular trichomes
C
fruit
D
non-glandular trichome
E
flowering branch
G
flower
I
gynoecium
K
seed
M
open corolla
N
fruiting calyx. Line drawing by P. Peralta; voucher A, C, K, N Nee 36164; B, D-J, L, M Eshbaugh 1943 b.
Figure 3.
Capsicum eximium
.
A, J
anthers in ventral and dorsal view respectively
B
flowering branch
C
flower
D, G
transverse section of the seed coat
E
fruit
F
embryo
H
gynoecium
I
seed
K
open corolla
L
seed in cross section
M
non-glandular trichome. Line drawing by N. Flury; voucher A, C-M: Hunziker 1907; B, Hunziker 7346.
Glandular hairs are rare in
Capsicum
. The dense indumentum of
Capsicum eshbaughii
consists of different types of glandular trichomes, some of them observed only in this species. The hair variations are: long simple hairs with multicellular and verrucate stalks and unicellular stipitate (
Fig. 2 F
) or not stipitate (
Fig. 2 B
) heads; short hairs with bicellular smooth stalks and multicellular heads (
Fig. 2 J
); and bifurcate hairs with both branches ending in a unicellular head (
Fig. 2 L
) or one branch non-glandular and the other longer and glandular (
Fig. 2 H
).
Capsicum eshbaughii
, together with
Capsicum eximium
and
Capsicum cardenasii
Heiser & P.G.Sm., is known as
"ulupica"
(Eshbaugh 1943 C); their very pungent fruits are very much appreciated as a spice or as pickles in the kitchens of Bolivia (
Eshbaugh and Smith 1971
).
This species is named in honor to Dr. W.H. Eshbaugh (Miami University) who first recognised the distinctness of this species and devoted part of his research to the taxonomy of chili peppers.
Specimens Examined
. Bolivia. Cochabamba:
Jose
Carrasco Torrico, camino Cochabamba-Comarapa, Copachuncho, ca. 3000 m, 29 Mar 1987, D.Flores 89 (CORD, LPB). Santa Cruz: Florida, El Sauce, west of Samaipata, elev. 1730 m, 30 Mar 1987, W.H.Eshbaugh 1943 C & D (CORD); same locality, 1 Aug 1990, W.H.Eshbaugh 1943 a (CORD); 5 km (by air) SE of Mairana, on road to Samaipata, at "Quebrada Seca", 18
°
09' S, 63
°
56' W, ca. 1550 m, 4 Feb 1988, M.Nee 36164 (CORD).