Transfer of Pygmaeotineacrisostomella Amsel, 1957 from Tineidae to Psychidae and its taxonomic status (Lepidoptera) Author Sobczyk, Thomas text Nota Lepidopterologica 2015 38 2 127 131 http://dx.doi.org/10.3897/nl.38.5090 journal article http://dx.doi.org/10.3897/nl.38.5090 2367-5365-2-127 9E0B96CCB4864B01849DD560DF146DFE Taxon classification Animalia Lepidoptera Psychidae Eumasia crisostomella Amsel, 1957 comb. n. Figs 1-6 Examined material. Holotype ♂ "vi [1]950 / Singeverga" back cover (handwritten Amsel) "Portugal / coll. Monteiro", "GU 3199", red label " Pygmaetinea crisostomella ", "coll. SMNK", genital slide with same information and additional "Holotypus" ; Paratype 1 ♀ "Singeverga / vi.1953" back cover (handwritten Amsel) "Portugal / coll. Monteiro", genital preparation 3198, red label "Allotypus ♀, leg. H. Amsel", "coll. SMNK". Citation of original description by Amsel. ( Pygmaeotinea ) Fuehler des ♂ kurz bewimpert, bis etwa 1/2 Costa reichend. Kopfhaare abstehend. Vflg.- Geaeder : Zelle offen, nur 3 Radialadern: r2+r3 und r3+r4 zu je einer Ader zusammengefallen; mit m1 und r4+r5 gestielt; m2 fehlend; ax ohne Wurzelschlinge. Hflg.- Geaeder : rr und m1 gestielt, die Spitze umgreifend, m2 fehlend. Axillaradern stark reduziert. Genitalapparat des ♂: Uncus abgerundet, Gnathos fehlend, Vinculum (Saccus) zugespitzt, Valven in einen basalen und einen terminalen Teil gegliedert, ohne sonstige Strukturen. (crisostomella) Spw. 7 mm. Kopfhaare gelblich, Palpen braeunlich . Fuehler des ♂ bis 1/2 Costa reichend, sehr kurz bewimpert, fast pubscent. Zwischen oberen Aussenrand und der Fuehlerwurzel ein grosser quecksilberartiger Fleck. Vfgl. Gelblich. Ein grosser braeunlicher Fleck am Innenrand bei 1/2 , weitere Flecke im apikalen Fluegelteil . Die Art ist durch den grossen , fast viereckigen Fleck am Innenrand bei 1/2 gut charakterisiert. An der Costa stehen einige braeunliche Schuppen, im aeusseren Drittel der Fluegel haeufen sich diese zu Flecken. Fransen graubraun. Hflg. Nur wenig schmaler als die Vflg. Fransen etwas laenger als der Fluegel breit ist. Beine ungeringelt. Genitalapparat des ♂: Uncus abgerundet, Gnathos fehlend. Vinculum ziemlich lang, zugespitzt. Valven aus zwei Teilen bestehend, einen aeusseren , nicht strukturierten und einen inneren, der deutlich gegenueber dem Aussenteil abgesetzt ist. Aedoeagus leicht gebogen, ziemlich lang, ohne Cornuti. Genitalpraeparat 3199 ( ... ) Einige Stuecke wurden aus abgeflachten Saecken gezuechtet , die 7 mm lang mit Sandkoernchen besetzt sind. Redescription crisostomella. Forewing length (with fringes) 3.5-4 mm, wingspan 7-8 mm. Labial palpi tripartite, median segment of double width and equal length as distal segment. Both segments covered with brown, ventrally directed hair-like scales. Antennal segments round, covered with short setae, dorsally with a semi-circular fan-like arrangement of broad, dark brown scales. The angle of this fan is 30-40° (see description of measurement by Haettenschwiler 1998 ). Forelegs with bristle brush (without epiphysis). Forewing scales long-oval, rounded distally, usually four- to six-pointed. Forewing ground colour yellowish, with a distinct dark brown spot at 1/2 of posterior margin and partly the anterior margin with some very narrow dark brown spots; distal third significantly spotted and converging partially to transverse lines. Fringe gray, hair-like, bi- or tricuspid. Hindwings uniformly dark gray. Male genitalia (genital preparation no. 3199 by Amsel): Total length 0.5 mm, very weakly sclerotized (perhaps an artifact due to excessive maceration). In ventral view, almost three times as long as wide. Valva extending beyond posterior margin of tegumen, weakly sclerotized distally with fine setae. Phallus 0.4 mm long, tubular, almost straight. Female (differences to males). Antenna filiform, wingspan 8 mm. Wing pattern as in the male, but missing dark spots at the anterior margin of basal 2/3 of the forewings. Abdominal segment eight with dense, wavy hairs. Female genitalia (genital preparation no. 3198 by Amsel): 2.0 mm total length. Oviscapt with fine setae distally, supported by three pairs of apophyses. Posterior apophyses with a length of 1.9 mm, almost as long as the entire genitalia, anterior apophyses 0.85 mm long, forked distally, distal edge of antevaginal plate concave, covered with fine, distally directed spines. Ovipositor also with a third pair of 0.5-mm-long apophyses. Figures 1-6. Eumasia crisostomella Amsel, 1957. 1. ♂ Holotype, Portugal, Singeverga, vi. [1]950. 2. Holotype genital preparation 3199. 3. Detail antennae. 4. Foreleg. 5. ♀ Paratype ♀ same data, but vi.1953. 6. ♀ Paratype genital preparation 3198. Scale bar: 1 mm (1, 5), 0.1 mm (2, 3), 0.5 mm (4, 6). Diagnosis. Eumasia crisostomella Amsel, 1957, comb. n. differs from Eumasia parietariella (Heydenreich, 1851) from Central and South Europe and Eumasia brunella Haettenschwiler , 1998 from the Iberian Peninsula by the forewing pattern. Forewings of the latter two species are also covered with dark spots basally, but the dark spots of Eumasia crisostomella are concentrated on distal third. On the basal 2/3 there are two or three small partial dark spots on the anterior margin. Most conspicuous is a large, almost square dark spot in the middle of the posterior margin. Hindwings of Eumasia parietariella are light cream, of Eumasia brunella brown and of Eumasia crisostomella grey. Remarks. The original description of Pygmaeotinea neither contains information about a differentiation from other genera nor why the description of a new genus was considered necessary. The venation (figure 5, Amsel 1957 ) and antennal and genital structures are clearly consistent with typical structures of Eumasia . A diagnosis for the genus is given by Haettenschwiler (1998) . Particularly noteworthy is the scalation of the male antennae with a dorsally semi-circular fan-like arrangement of broad scales. Each antennal segment has a dorsal compartment of broader scales. Thus, Pygmaeotinea Amsel, 1957, syn. n. is identical to Eumasia Chretien , 1904 and is synonymized here with it and its type species transferred to Eumasia , as Eumasia crisostomella Amsel, 1957, comb. n. The label data do not completely match with information provided in the original description. Holotype ♂ denotes VI. [1]950 as date. There was no label on the pinned specimen that indicated it as the holotype. That information was only present on the genital slide belonging to this specimen and the slide number 3199 is present on both, the genitalia slide and on the pinned holotype specimen. In the description the date for the female (allotype) is listed as vi.1950; however, the specimen label reads 'vi.1953' .