Synopsis of the species of Cheilolejeunea (Marchantiophyta, Lejeuneaceae) in the Pacific dominion and Páramo province of tropical America
Author
Gil-Novoa, Jorge Enrique
Programa de Pós-graduação em Botânica, Escola Nacional de Botânica Tropical, Instituto de Pesquisas Jardim Botânico do Rio de Janeiro. Rua Pacheco Leão, Horto, 2040, 22460 - 080, Rio de Janeiro-RJ, Brazil. & Grupo Sistemática Biológica, Herbario UPTC, Universidad Pedagógica y Tecnológica de Colombia, Tunja, Boyacá, Colombia.
Author
Costa, Denise Pinheiro
0000-0001-9495-3029
Instituto de Pesquisas Jardim Botânico do Rio de Janeiro - IPJBRJ, Rua Pacheco Leão 915, 22460 - 030, Rio de Janeiro, RJ, Brazil. denisepinheirodacosta @ gmail. com; https: // orcid. org / 0000 - 0001 - 9495 - 3029
denisepinheirodacosta@gmail.com
text
Phytotaxa
2023
2023-03-14
587
2
73
120
http://dx.doi.org/10.11646/phytotaxa.587.2.1
journal article
10.11646/phytotaxa.587.2.1
1179-3163
7731813
10.
Cheilolejeunea erostrata
Schuster (1978: 427)
.
Type:—
VENEZUELA
.
Tachira
:
Páramo
de Tama
, base of
Pico
de Banderas
,
3140 m
,
1 March 1976
,
R
.
M
.
Schuster & L.
Ruíz-Terán 76-1921e
(
holotype
F
!)
.
Plants yellowish green to light brown,
1–1.6 mm
wide. Ventral merophytes 2 cells wide. Leaves imbricate, orbicular to ovate-elongate, recurved at the apex, 680–950 × 400–650 µm; margin entire; apex acute to more commonly apiculate; dorsal and ventral margin slightly recurved; cells orbicular to ovate, mammillose to papillose on the dorsal side, conspicuous trigones; basal cells 20–30 × 15–23 µm, median cells 16–28 × 14–22 µm, marginal cells 10–17 × 8–16 µm. Lobules rectangular, 1/3–1/2 of leaf length, free margin slightly involute, tooth very short (1 cell long), observed in few lobules; keel slightly arched. Underleaves subimbricate to commonly imbricate, widely orbicular to obovate, 450– 500 × 470–530 µm, 5 x stem width, bifid to 1/5–1/3, with a V-shaped sinus, margin entire, insertion strongly curved. Autoicous. Androecia commonly on the main stem, 3–4 pairs of orbicular bracts. Gynoecia, with pycnolejeuneoid innovations, bracts obovate, apex acute. Perianth 5-keeled, keels weak to very strong, without beak.
Distribution and habitat
:—Endemic to the Venezuelan Andean páramos, on the border with
Colombia
; this species is only known by the
type
collection, growing on the branches of
Scrophulariaceae
shrubs, in grass páramo between
3100–3500 m
, in the Páramo province (
Fig. 8
).
Notes
:—The species is characterized by the large plants with large, apparently flat and rectangular lobules (1/3– 1/2 of leaf length) but with a slightly involute free margin and very short lobule teeth (1 cell long), and perianths without a beak. More exploration is needed of the páramos between
Colombia
and
Venezuela
, to confirm the continued occurrence of the species in the area.
Taking into account its extent of occurrence and number of locations (
Hallingbäck 1998
, UCN 2001), it is considered here as Critically Endangered (CR-B1a).